The Northumberland Formation is a Late Cretaceous (?Campanian-?Maastrichtian)-aged geologic formation in Canada. It belongs to the larger Nanaimo Group. Indeterminate bird and pterosaur fossils have been recovered from the formation,[1] as well as a potential gladius of Eromangateuthis.[2] An extensive diversity of shark teeth is known from the formation; many appear to be closely allied with modern deep-water shark taxa, suggesting a deep-water environment for the formation.[3] The most well-known exposures of the formation are on Hornby Island.
Northumberland Formation | |
---|---|
Stratigraphic range: Late Cretaceous, ? | |
Type | Geological formation |
Unit of | Nanaimo Group |
Underlies | Spray Formation |
Location | |
Country | Canada |
Genus | Species | Location | Abundance | Notes | Images |
---|---|---|---|---|---|
Meristodonoides | M. sp. | A hybodontid shark. |
Genus | Species | Location | Abundance | Notes | Images |
---|---|---|---|---|---|
Chlamydoselachus | C. balli | A chlamydoselachid shark, related to the modern frilled shark. | |||
Rolfodon | R. ludvigseni | A chlamydoselachid shark. | |||
R. cf. thompsoni | |||||
Dykeius | D. garethi | A very large chlamydoselachid shark. | |||
Hexanchus | H. microdon | A cow shark related to the modern bluntnose sixgill shark. | |||
Notidanodon | N. pectinatus | A cow shark. | |||
Xampylodon | X. dentatus | A large cow shark. | |||
Protoheptranchias | P. lowei | A cow shark similar to the modern sharpnose sevengill shark. | |||
Paraorthacodus | P. rossi | A paraorthacodontid shark. | |||
Komoksodon | K. kwutchakuth | A komoksodontid shark. |
Genus | Species | Location | Abundance | Notes | Images |
---|---|---|---|---|---|
Echinorhinus | E. lapaoi | A echinorhinid shark, related to the modern bramble shark. |
Genus | Species | Location | Abundance | Notes | Images |
---|---|---|---|---|---|
Eoetmopterus | E. supercretaceus | An etmopterid shark similar to modern lantern sharks. | |||
Squalus | S. vondermarcki | A dogfish shark, related to modern spurdogs. | |||
S. nicholsae | |||||
S. sp. | |||||
Centrosqualus | C. mustardi | A dogfish shark. | |||
Protocentrophorus | P. steviae | A dogfish shark. | |||
Rhinoscymnus | R. clarki | A sleeper shark, possibly included within Somniosus. | |||
Centroscymnus | C. sp. | A sleeper shark related to the modern Portuguese dogfish. | |||
Squaliodalatias | S. savoiei | A dalatiid shark. | |||
Hessinodon | H. wardi | A possibly dalatiid shark. |
Genus | Species | Location | Abundance | Notes | Images |
---|---|---|---|---|---|
Pristiophorus | P. smithi | A sawshark. | |||
P. pricei |
Genus | Species | Location | Abundance | Notes | Images |
---|---|---|---|---|---|
Plicatoscyllium | P. cf. globidens | A ginglymostomatid shark. | |||
Hemiscyllium | H. hermani | A bamboo shark. |
Genus | Species | Location | Abundance | Notes | Images |
---|---|---|---|---|---|
Carcharias | C. dominguei | A sand shark, related to the modern sand tiger shark. |
Genus | Species | Location | Abundance | Notes | Images |
---|---|---|---|---|---|
Synechodus | S. dereki | A palaeospinacid shark. |
Genus | Species | Location | Abundance | Notes | Images |
---|---|---|---|---|---|
Scyliorhinidae indent. | A catshark of uncertain affinities. | ||||
Florenceodon | F. johnyi | A florenceodontid shark. |
Genus | Species | Location | Abundance | Notes | Images |
---|---|---|---|---|---|
Gwawinapterus | G. beardi | Jaw. | A saurodontid ichthyodectiform, initially identified as an istiodactylid pterosaur.[5] |
Genus | Species | Location | Abundance | Notes | Images |
---|---|---|---|---|---|
Maaqwi[6] | M. cascadensis | Coracoid and wing bones. | A large vegaviid ornithuran. | ||
Enantiornithes indet.[7] | An enanthiornithine avialan. |
Genus | Species | Location | Abundance | Notes | Images |
---|---|---|---|---|---|
Azhdarchoidea indet.[1] | A possibly azhdarchid pterosaur. |
Genus | Species | Location | Abundance | Notes | Images |
---|---|---|---|---|---|
Mosasauridae indet.[8] | A mosasaur. |
Genus | Species | Location | Abundance | Notes | Images |
---|---|---|---|---|---|
Baculites | B. occidentalis | A baculitid ammonite. | |||
Fresvillia | F. constricta | A baculitid ammonite. | |||
Diplomoceras | D. cylindraceum | A diplomoceratid ammonite. | |||
D. cf. cylindraceum | |||||
Exiteloceras | E. densicostatum | A diplomoceratid ammonite. | |||
E. bipunctatum | |||||
Phylloptychoceras | P. horitai | A diplomoceratid ammonite. | |||
Solenoceras | S. exornatus | A diplomoceratid ammonite. | |||
S. cf. reesidei | |||||
Nostoceras | N. adrotans | A nostoceratid ammonite. | |||
N. hornbyensis | |||||
N. aff. pauper | |||||
Enchoteuthis[2] | E. sp. | A muensterellid octopodiform. | |||
Eromangateuthis[2] | E. soniae? | A plesioteuthid octopodiform. | |||
Cyrtobelus[10] | C. hornbyense | A groenlandibelid spirulid.[2] | |||
Actinosepia[2] | A. canadensis | An actinosepiid vampyromorphid.[11] | |||
Cirroteuthidae indet.[2] | Gladius. | A cirroteuthid octopus. |
Genus | Species | Location | Abundance | Notes | Images |
---|---|---|---|---|---|
Tessarolax[8] | T. louellae | An aporrhaid gastropod. |
Genus | Species | Location | Abundance | Notes | Images |
---|---|---|---|---|---|
Neocallichirus | N. manningi | A callianassid decapod. | |||
Longusorbis | L. cuniculosus | A longusorbiid decapod. | |||
Unnamed raninoid[12] | Unnamed | A raninoid decapod. |
Genus | Species | Location | Abundance | Notes | Images |
---|---|---|---|---|---|
Cunninghamia | C. hornbyensis | Permineralized wigs and leaves. | A member of the extant Cupressaceae genus Cunninghamia. Closely resembles extant species.[13] | ||
Cycadeoidea[14] | C. maccafferyii | A bennettitalean. |
Genus | Species | Location | Abundance | Notes | Images |
---|---|---|---|---|---|
Atli[15] | A. morinii | Stem | A member of the Ranunculales with a liana-like growth habit. |
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