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2016 in paleontology

2016 in paleontology

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Paleontology or palaeontology (from Greek: paleo, "ancient"; ontos, "being"; and logos, "knowledge") is the study of prehistoric life forms on Earth through the examination of plant and animal fossils.[1] This includes the study of body fossils, tracks (ichnites), burrows, cast-off parts, fossilised feces (coprolites), palynomorphs and chemical residues. Because humans have encountered fossils for millennia, paleontology has a long history both before and after becoming formalized as a science. This article records significant discoveries and events related to paleontology that occurred or were published in the year 2016.

Plants

Cnidarians

Research

New taxa

Name Novelty Status Authors Age Unit Location Notes Images

Aphyllum pachystele[4]

Sp. nov

Valid

Munson & Jell

Silurian (late Wenlock)

Jack Formation

 Australia

A rugose coral belonging to the family Tryplasmatidae.

Aulokoninckophyllum potyi[5]

Sp. nov

Valid

Somerville, Rodríguez & Said

Carboniferous

Azrou-Khenifra Basin

 Morocco

An aulate coral.

Axuolites higoensis[6]

Sp. nov

Valid

Niko

Silurian (Ludlow)

Fukami Formation

 Japan

A tabulate coral belonging to the order Favositida and the family Coenitidae.

Bjarmia[7]

Gen. et sp. nov

Valid

Grazhdankin

Late Ediacaran (~550 Ma)

Erga Formation

 Russia

A jellyfish-like organism, a possible relative of scyphomedusae. Genus includes new species B. cycloplerusa.

Byronia jaegeri[8]

Sp. nov

Valid

Vinn et al.

Ordovician (Katian)

 Estonia

A possible thecate scyphozoan.

Calceola collini[9]

Sp. nov

Valid

Wright, Plusquellec & Gourvennec

Devonian (Eifelian)

Saint-Fiacre Formation

 France

An operculate coral belonging to the family Calceolidae; a species of Calceola.

Cambroctoconus coreaensis[10]

Sp. nov

Valid

Park et al.

Cambrian

Daegi Formation

 South Korea

A stem-cnidarian.

Caninophyllum charli[11]

Sp. nov

Valid

Denayer

Carboniferous (Tournaisian)

Yılanlı Formation

 Turkey

A rugose coral belonging to the family Bothrophyllidae; a species of Caninophyllum.

Cystodactylon[12]

Gen. et sp. nov

Valid

Denayer & Webb

Carboniferous (Tournaisian)

Gudman Formation

 Australia

A rugose coral. Genus includes new species C. orbum.

Dokophyllum hillae[4]

Sp. nov

Valid

Munson & Jell

Silurian (Ludfordian)

Jack Formation

 Australia

A rugose coral belonging to the family Ketophyllidae.

Edaphophyllum zhaotongense[13]

Sp. nov

Valid

Liao & Ma

Devonian (Emsian)

 China

A rugose coral.

Eoaequorea[14]

Gen. et sp. nov

Valid

Tang et al.

Ediacaran

Doushantuo Formation

 China

A member of Hydrozoa of uncertain phylogenetic placement. The type species is Eoaequorea xingi.

Eosiderastrea[15]

Gen. et comb. et 2 sp. nov

Valid

Löser

Cretaceous (late Valanginian to Santonian)

 Austria
 France
 Germany
 Greece
 Iran
 Italy
 Japan
 Mexico
 Poland
 Spain
 Ukraine
 United States
 Uzbekistan

A stony coral belonging to the family Siderastreidae. The type species is "Siderastrea" cuyleri Wells (1932); genus also includes "Phyllocoenia" cotteaui Orbigny (1850), "Diploastrea" crassa Kuzmicheva (1980), "Diploastrea" crassicostata Morycowa & Masse (1998), "Phyllocoenia" cyclops Felix (1891), "Confusastrea" dollfusi Prever (1909), "Confusastrea" felixi Prever (1909); "Stephanocoenia" grandipora Orbigny (1849), "Diploastrea" harrisi Wells (1932), "Diploastrea" hilli Wells (1933), "Montastraea" nagaoi Eguchi (1951), "Pleurocora" reussi Milne Edwards (1857), "Diplocoenia" splendida Prever (1909), "Diploastrea" vaughani Wells (1933) and "Plesiofavia" villaltai Reig Oriol (1991), as well as new species E. paragrandipora Löser (2016) and E. stefani Löser (2016).[16]

Famastraea[17]

Gen. et sp. nov

Valid

Berkowski, Zapalski & Wrzołek

Devonian (Famennian)

 Poland

A coral belonging to the group Rugosa and the family Campophyllidae. The type species is Famastraea catenata.

Fomichevella najafi[18]

Sp. nov

Valid

Badpa et al.

Carboniferous (Bashkirian)

Sardar Formation

 Iran

A member of Rugosa belonging to the group Stauriida and the family Kleopatrinidae.

Gertholites haikawai[19]

Sp. nov

Valid

Niko & Fujikawa

Early Carboniferous

Akiyoshi Limestone Group

 Japan

A tabulate coral.

Gerviphyllum[9]

Gen. et comb. nov

Valid

Wright, Plusquellec & Gourvennec

Early Devonian (likely late Lochkovian to early Pragian)

 France

An operculate coral belonging to the family Calceolidae; a new genus for "Calceola" gervillei Bayle (1878).

Gudmania[12]

Gen. et sp. nov

Valid

Denayer & Webb

Carboniferous (Tournaisian)

Gudman Formation

 Australia

A rugose coral. Genus includes new species G. darumbalae.

Heintzella fluegeli[18]

Sp. nov

Valid

Badpa et al.

Carboniferous (Bashkirian)

Sardar Formation

 Iran

A member of Rugosa belonging to the group Stauriida and the family Kleopatrinidae.

Kullingia rotadiscopsis[14]

Sp. nov

Valid

Tang et al.

Ediacaran

Doushantuo Formation

 China

A member of Hydrozoa of uncertain phylogenetic placement; a species of Kullingia.

Madrepora mitsukurii[20]

Sp. nov

Valid

Niko, Suzuki & Taguchi

Miocene

Katsuta Group

 Japan

A madrepore.

Marekostragulum grafi[21]

Sp. nov

Valid

Plusquellec, Galle & Franke

Devonian (Emsian)

 Germany

A tabulate coral belonging to the family Hyostragulidae.

Mesoconularia (?) lukesi[22]

Sp. nov

Valid

Mergl, Ferrova & Frýda

Devonian (late Emsian)

Suchomasty Limestone

 Czech Republic

A member of Conulariida, possibly a species of Mesoconularia.

?Michelinia vinni[23]

Sp. nov

Valid

Zapalski, Berkowski & Wrzołek

Devonian (late Famennian)

 Poland

A tabulate coral belonging to the family Micheliniidae; possibly a species of Michelinia.

Multicarinophyllum vepreculatum[4]

Sp. nov

Valid

Munson & Jell

Silurian (Ludfordian)

Jack Formation

 Australia

A rugose coral belonging to the family Amsdenoididae.

Oligophylloides maroccanus[24]

Sp. nov

Valid

Weyer

Devonian (late Famennian)

 Morocco

A member of Heterocorallia.

Palaeosiderofungia[15]

Gen. et comb. nov

Valid

Löser

Cretaceous (Aptian to Santonian)

 Austria
 Czech Republic
 France
 Germany
 Greece
 Serbia

A stony coral belonging to the family Siderastreidae; a new genus for "Thamnasteria" exigua Reuss (1854).

Parostragulum[21]

Gen. et sp. nov

Valid

Plusquellec, Galle & Franke

Devonian (Emsian)

 Germany

A tabulate coral belonging to the family Hyostragulidae. The type species is P. problematicum.

Pleurodictyum goldfussi[25]

Sp. nov

Valid

Plusquellec & Franke

Devonian (Emsian)

Wiltz Beds

 Germany
 Luxembourg

A tabulate coral belonging to group Favositida and to the family Micheliniidae.

Pleurodictyum pruemensis[25]

Sp. nov

Valid

Plusquellec & Franke

Devonian (Emsian)

Wiltz Beds

 Germany

A tabulate coral belonging to group Favositida and to the family Micheliniidae.

Potyphyllum[26]

Gen. et comb. nov

Valid

Coen-Aubert

Devonian (late Frasnian)

 Belgium
 Czech Republic
 France
 Germany
 Poland
 Russia
 United Kingdom

A rugose coral belonging to the family Phillipsastreidae. The type species is "Cyathophyllum" ananas Goldfuss (1826); genus also includes "Pseudoacervularia" dybowskii Różkowska (1953), "Pseudoacervularia" intercellulosa (Phillips, 1841) sensu Pickett (1967), "Schlüteria" lyskovensis Ermakova (1957), "Acervularia" macrommata Roemer (1855), "Phillipsastrea" plantana Różkowska (1979), "Cyathophyllum" profundum Michelin (1845), "Phillipsastrea" rozkowskae Scrutton (1968), "Pseudoacervularia" cf. smithi (Różkowska, 1953) sensu Pickett (1967), "Phillipsastrea" ananas veserensis Coen-Aubert (1974) and "Phillipsastrea" zerda Galle, 1992 in Hladil et al. (1992).

Preisingerella[27]

Gen. et sp. nov

Valid

Kossovaya, Novak & Weyer

Early Permian

 Slovenia

A coral similar to members of the genus Caninia. The type species is P. stegovnikensis.

Ptychophyllum variatum[4]

Sp. nov

Valid

Munson & Jell

Silurian (late Gorstian to Ludfordian)

Jack Formation

 Australia

A rugose coral belonging to the family Ptychophyllidae.

Pycnostylus polyphyllodus[4]

Sp. nov

Valid

Munson & Jell

Silurian (late Wenlock)

Jack Formation

 Australia

A rugose coral belonging to the family Pycnostylidae.

Sokolovia[28]

Gen. et sp. nov

Junior homonym

Tsyganko

Late Devonian (Famennian)

 Russia
( Komi Republic)

A tabulate coral belonging to the family Pachyporidae. The type species is Sokolovia pershinae. The generic name is preoccupied by Sokolovia Ilovaisky (1934) and Sokolovia Shishkinskaya (1964).

Spinaxon[29]

Gen. et sp. nov

Valid

Weyer

Devonian (late Frasnian)

 France
 Germany

A member of Rugosa belonging to the family Petraiidae. The type species is Spinaxon potyi.

Syringopora hilarowiczi[23]

Sp. nov

Valid

Zapalski, Berkowski & Wrzołek

Devonian (late Famennian)

 Poland

A tabulate coral belonging to the family Syringoporidae; a species of Syringopora.

Syringopora kowalensis[23]

Sp. nov

Valid

Zapalski, Berkowski & Wrzołek

Devonian (late Famennian)

 Poland

A tabulate coral belonging to the family Syringoporidae; a species of Syringopora.

Talfania[30]

Gen. et sp. nov

Valid

Peel & McDermott

Ordovician (Katian)

Sholeshook Limestone Formation

 United Kingdom

A solitary coral. The type species is Talfania calicula.

Thamnoptychia mistiaeni[23]

Sp. nov

Valid

Zapalski, Berkowski & Wrzołek

Devonian (late Famennian)

 Poland

A tabulate coral belonging to the family Pachyporidae; a species of Thamnoptychia.

Utaratuia yunnanensis[13]

Sp. nov

Valid

Liao & Ma

Devonian (Eifelian)

 China

A rugose coral.

Vassiljukia[31]

Gen. et comb. nov

Valid

Denayer & Ogar

Carboniferous (Viséan)

 Turkey
 Ukraine

A colonial rugose coral; a new genus for "Lithostrotion" columnariformis Vassiljuk (1960).

Vesicospina[4]

Gen. et sp. nov

Valid

Munson & Jell

Silurian (Ludfordian)

Jack Formation

 Australia

A rugose coral belonging to the family Kyphophyllidae. The type species is V. julli.

Vulykhia[32]

Nom. nov

Valid

Doweld

Late Ordovician

 Mongolia

An anthozoan belonging to the superfamily Heliolitoidea; a replacement name for Concavites Bondarenko & Minzhin (1981).

Arthropods

Bryozoans

Name Novelty Status Authors Age Unit Location Notes Images

Anisotrypa kjarkiensis[33]

Sp. nov

Valid

Tolokonnikova

Carboniferous (Tournaisian)

 Armenia

A bryozoan belonging to the group Trepostomata and the family Anisotrypidae.

Beisselina skyscanica[34]

Sp. nov

Valid

Koromyslova & Pakhnevich

Late Cretaceous (Campanian)

 Belarus

A bryozoan belonging to the group Ascophora and the family Tessaradomidae; a species of Beisselina.

Celleporaria pirabasensis[35]

Sp. nov

Valid

Muricy et al.

Oligocene-Miocene

Pirabas Formation

 Brazil

A bryozoan belonging to the group Cheilostomata and the family Lepraliellidae.

Celleporaria triangulavicularis[35]

Sp. nov

Valid

Muricy et al.

Oligocene-Miocene

Pirabas Formation

 Brazil

A bryozoan belonging to the group Cheilostomata and the family Lepraliellidae.

Dybowskiella hupehensiformis[36]

Sp. nov

Valid

Ernst

Permian

Zongba Formation

 China

A member of Cystoporata belonging to the family Fistuliporidae.

Dyscritella lii[36]

Sp. nov

Valid

Ernst

Permian

Zongba Formation

 China

A member of Trepostomata belonging to the family Dystritellidae.

Ehrhardina[37]

Gen. et 2 sp. nov

Valid[38]

Martha & Taylor

Late Cretaceous (Cenomanian)

 France
 Germany
 United Kingdom

A bryozoan belonging to the group Flustrina and the family Onychocellidae. The type species is Ehrhardina voigti; genus also includes Ehrhardina pikeae.

Eridotrypella danzikensis[39]

Sp. nov

Valid

Tolokonnikova

Devonian (Famennian)

 Azerbaijan

A bryozoan belonging to the group Trepostomata and the family Eridotrypellidae.

Etherella tibetensis[36]

Sp. nov

Valid

Ernst

Permian

Zongba Formation

 China

A member of Cystoporata belonging to the family Etherellidae.

Fistuliphragma moniliformis[40]

Sp. nov

Valid

Mesentseva

Devonian (Emsian)

 Russia

A bryozoan belonging to the group Cystoporida.

Fistuliphragma sibirica[40]

Sp. nov

Valid

Mesentseva

Devonian (Emsian)

 Russia

A bryozoan belonging to the group Cystoporida.

Fistulipora sakagamii[36]

Sp. nov

Valid

Ernst

Permian

Zongba Formation

 China
 Thailand

A member of Cystoporata belonging to the family Fistuliporidae.

Fistulipora salairiensis[40]

Sp. nov

Valid

Mesentseva

Devonian (Emsian)

 Russia

A bryozoan belonging to the group Cystoporida.

Fistuliramus fasciculus[40]

Sp. nov

Valid

Mesentseva

Devonian (Emsian)

 Russia

A bryozoan belonging to the group Cystoporida.

Fistulocladia cincinnata[40]

Sp. nov

Valid

Mesentseva

Devonian (Emsian)

 Russia

A bryozoan belonging to the group Cystoporida.

Margaretta amplipora[41]

Sp. nov

Valid

Sonar & Gaikwad

Cenozoic

 India

An ascophoran belonging to the family Margarettidae.

Margaretta guhai[41]

Nom. nov

Valid

Sonar & Gaikwad

Cenozoic

 India

An ascophoran belonging to the family Margarettidae.

Margaretta hariparensis[41]

Sp. nov

Valid

Sonar & Gaikwad

Cenozoic

 India

An ascophoran belonging to the family Margarettidae.

Metastenodiscus[42]

Gen. et comb. nov

Valid

Ernst, Schäfer & Grant-Mackie

Late Triassic

 New Caledonia
 New Zealand

A trepostome bryozoan. A new genus for "Stenodiscus" zealandicus Schäfer & Grant-Mackie (1994) and "Stenodiscus" kawhiae Schäfer & Grant-Mackie (1994).

Nikiforopora arpaensis[33]

Sp. nov

Valid

Tolokonnikova

Carboniferous (Tournaisian)

 Azerbaijan

A bryozoan belonging to the group Trepostomata and the family Stenoporidae.

Pachydermopora grodnoensis[34]

Sp. nov

Valid

Koromyslova & Pakhnevich

Late Cretaceous (Campanian)

 Belarus

A bryozoan belonging to the group Ascophora and the family Tessaradomidae; a species of Pachydermopora.

Petalotrypa myunkhbalaensis[39]

Sp. nov

Valid

Tolokonnikova

Devonian (Famennian)

 Azerbaijan

A bryozoan belonging to the group Trepostomata.

Planicellaria walsariensis[43]

Sp. nov

Valid

Sonar & Pawar

Cenozoic

 India

A bryozoan belonging to the group Cheilostomata and the family Calloporidae.

Protoretepora irregularis[36]

Sp. nov

Valid

Ernst

Permian

Zongba Formation

 China

A member of Fenestrata belonging to the family Polyporidae.

Pyriporella charopadiensis[43]

Sp. nov

Valid

Sonar & Pawar

Cenozoic

 India

A bryozoan belonging to the group Cheilostomata and the family Calloporidae.

Pyriporella vadsariensis[43]

Sp. nov

Valid

Sonar & Pawar

Cenozoic

 India

A bryozoan belonging to the group Cheilostomata and the family Calloporidae.

Rectifenestella famenniensis[39]

Sp. nov

Valid

Tolokonnikova

Devonian (Famennian)

 Azerbaijan

A bryozoan belonging to the family Fenestellidae.

Rectifenestella kadrluiensis[39]

Sp. nov

Valid

Tolokonnikova

Devonian (Famennian)

 Armenia

A bryozoan belonging to the family Fenestellidae.

Stenophragmidium buckhornensis[44]

Sp. nov

Valid

Ernst et al.

Carboniferous (Pennsylvanian)

Boggy Formation

 United States
( Oklahoma)

Streblotrypa (Streblotrypa) heltzelae[44]

Sp. nov

Valid

Ernst et al.

Carboniferous (Pennsylvanian)

Boggy Formation

 United States
( Oklahoma)

Streblotrypa (Streblotrypa) parviformis[36]

Sp. nov

Valid

Ernst

Permian

Zongba Formation

 China

A member of Cryptostomata belonging to the family Hyphasmoporidae.

Tabuliporella nakhichevanica[33]

Sp. nov

Valid

Tolokonnikova

Carboniferous (Tournaisian)

 Azerbaijan

A bryozoan belonging to the group Trepostomata and the family Crustoporidae.

Tibetiporella[36]

Gen. et sp. nov

Valid

Ernst

Permian

Zongba Formation

 China

A member of Fenestrata belonging to the family Polyporidae. The type species is T. ornata.

Timanotrypa australis[36]

Sp. nov

Valid

Ernst

Permian

Noonkanbah Formation
Rat Buri Limestone
Zongba Formation

 Australia
 China
 Thailand

A member of Cryptostomata belonging to the family Timanodictyidae.

Zigzagopora[45]

Gen. et sp. nov

Valid

Wilson & Taylor

Ordovician (Sandbian)

Bromide Formation

 United States
( Oklahoma)

A cyclostome bryozoan. Genus includes new species Z. wigleyensis.

Brachiopods

Name Novelty Status Authors Age Unit Location Notes Images

Acritosia ogamensis[46]

Sp. nov

Valid

Tazawa et al.

Early Permian (Kungurian)

Nabeyama Formation

 Japan

Acrosaccus scutatus[47]

Sp. nov

Valid

Percival in Percival et al.

Ordovician

 Australia

A member of Discinidae.

Anathyris (Anathyris) calestiennensis[48]

Sp. nov

Valid

Mottequin et al.

Devonian (Frasnian)

Nismes Formation

 Belgium

Anisopleurella antiqua[49]

Sp. nov

Valid

Popov, Kebriaee-Zadeh & Pour

Ordovician (Darriwilian)

Lashkarak Formation

 Iran

A member of Strophomenida belonging to the family Sowerbyellidae.

Apatobolus anoskelidion[47]

Sp. nov

Valid

Percival in Percival et al.

Ordovician

 Australia

A member of Obolidae.

Atansoria australis[47]

Sp. nov

Valid

Percival in Percival et al.

Ordovician

 Australia

A member of Obolidae.

Atryparia (Costatrypa) agricolae[50]

Sp. nov

Valid

Halamski & Baliński in Baliński, Racki & Halamski

Devonian (Frasnian)

 Poland

A member of Atrypidae.

Aulacothyris maendlii[51]

Sp. nov

Valid

Sulser

Middle Jurassic (Callovian)

  Switzerland

A member of Terebratulida belonging to the family Zeilleriidae.

Aulacothyris waikatoensis[52]

Sp. nov

Valid

MacFarlan

Jurassic

 New Zealand

A member of Terebratulida.

Bellimurina fluctuosa[49]

Sp. nov

Valid

Popov, Kebriaee-Zadeh & Pour

Ordovician (Darriwilian)

Lashkarak Formation

 Iran

A member of Strophomenida belonging to the family Strophomenidae.

Biernatia pseudoplana[47]

Sp. nov

Valid

Engelbretsen in Percival et al.

Ordovician

 Australia

A member of Lingulata belonging to the family Biernatiidae.

Biernatia wrighti[47]

Sp. nov

Valid

Engelbretsen in Percival et al.

Ordovician

 Australia

A member of Lingulata belonging to the family Biernatiidae.

Biernatium minus[50]

Sp. nov

Valid

Baliński in Baliński, Racki & Halamski

Devonian (Frasnian)

 Poland

A member of Mystrophoridae.

Buxtonia inexpletucosta[53]

Sp. nov

Valid

Torres-Martínez & Sour-Tovar

Carboniferous (middle Pennsylvanian)

Ixtaltepec Formation

 Mexico

A member of Productoidea.

Canalilatus musculosus[54]

Sp. nov

Valid

Percival, Engelbretsen & Peng

Cambrian

Huaqiao Formation

 China

A lingulate brachiopod belonging to the family Zhanatellidae.

Cerasinella[55]

Nom. nov

Valid

Copper

Silurian (Llandovery)

Merrimack Formation

 Canada
( Quebec)

An atrypoid brachiopod; a replacement name for Cerasina Copper (1995) (preoccupied).

Chapinella belkovskensis[56]

Sp. nov

Valid

Baranov, Sokiran & Blodgett

Devonian (Famennian)

Nerpalakhsk Formation

 Russia
( Sakha Republic)

A member of Rhynchonellida belonging to the family Pugnacidae.

Chilcatreta[57]

Gen. et sp. nov

Valid

Lavié & Benedetto

Ordovician (Darriwilian)

San Juan Formation

 Argentina

A siphonotretid brachiopod. Genus includes new species C. tubulata.

Crispithyris[52]

Gen. et sp. nov

Valid

MacFarlan

Jurassic

 New Zealand

A member of Terebratulida. Genus includes new species C. nauarchus.

Cyrtina transcaucasica[58]

Sp. nov

Valid

Gretchishnikova & Oleneva in Oleneva

Devonian (Givetian)

 Azerbaijan

A member of Spiriferinida belonging to the family Cyrtinidae.

Davidsonia enmerkaris[50]

Sp. nov

Valid

Halamski in Baliński, Racki & Halamski

Devonian (Frasnian)

 Poland

A member of Davidsoniidae.

Dictyoclostus transversum[53]

Sp. nov

Valid

Torres-Martínez & Sour-Tovar

Carboniferous (middle Pennsylvanian)

Ixtaltepec Formation

 Mexico

A member of Productoidea.

Disculina mancenidoi[52]

Sp. nov

Valid

MacFarlan

Jurassic

 New Zealand

A member of Terebratulida.

Dolerorthis nadruvensis[59]

Sp. nov

Valid

Paškevičius & Hints

Ordovician (Katian)

 Lithuania

A member of Orthida belonging to the family Hesperorthidae.

Dulankarella hyrcanica[49]

Sp. nov

Valid

Popov, Kebriaee-Zadeh & Pour

Ordovician (Darriwilian)

Lashkarak Formation

 Iran

A member of Strophomenida belonging to the family Leptellinidae.

Dyoros (Lissosia) maya[60]

Sp. nov

Valid

Torres-Martínez, Sour-Tovar & Barragán

Permian (Leonardian)

Paso Hondo Formation

 Mexico

Echinocoelia parva[50]

Sp. nov

Valid

Baliński in Baliński, Racki & Halamski

Devonian (Frasnian)

 Poland

A member of Ambocoeliidae.

Eochonetes maearum[61]

Sp. nov

Valid

Bauer & Stigall

Late Ordovician

 United States
( Wyoming)

Eochonetes minerva[61]

Sp. nov

Valid

Bauer & Stigall

Late Ordovician

 United States
( Texas)

Eochonetes voldemortus[61]

Sp. nov

Valid

Bauer & Stigall

Late Ordovician

Saturday Mountain Formation

 United States
( Idaho)

Eoconulus puteus[47]

Sp. nov

Valid

Engelbretsen in Percival et al.

Ordovician

 Australia

A member of Lingulata belonging to the family Eoconulidae.

Eolingularia[62]

Gen. et comb. nov

Valid

Bitner & Emig

Carboniferous to Triassic

 China
 Russia
 Spain

A member of Lingulata belonging to the group Lingulida and the family Lingulidae. The type species is "Lingularia" siberica Biernat & Emig (1993).

Experilingula larga[54]

Sp. nov

Valid

Percival, Engelbretsen & Peng

Cambrian

Huaqiao Formation

 China

A member of Obolidae.

Flexaria magna[53]

Sp. nov

Valid

Torres-Martínez & Sour-Tovar

Carboniferous (middle Pennsylvanian)

Ixtaltepec Formation

 Mexico

A member of Productoidea.

Globiella kamiyassensis[63]

Sp. nov

Valid

Tazawa

Permian (Wordian)

 Japan

Glossella cuyanica[57]

Sp. nov

Valid

Lavié & Benedetto

Ordovician (Darriwilian)

San Juan Formation

 Argentina

Gondwanorthis[64]

Gen. et comb. nov

Valid[65]

Benedetto & Muñoz

Early Ordovician

 Argentina
 Iran

A plectorthoid brachiopod. A new genus for "Nanorthis" calderensis Benedetto (2007); genus also includes "Nanorthis" bastamensis Ghobadi Pour, Kebriaee-Zadeh & Popov (2011).

Gowanella[66]

Gen. et sp. nov

Valid

Hiller

Late Cretaceous (probably Maastrichtian)

Broken River Formation

 New Zealand

A member of Terebratulida related to Ostreathyris. The type species is G. capralis.

Grandispirifer qaidamensis[67]

Sp. nov

Valid

Lee, Shi & Chen in Shi et al.

Carboniferous (Serpukhovian)

Huaitoulata Formation

 China

A member of Spiriferoidea belonging to the family Spiriferidae.

Gretchispirifer[58]

Gen. et sp. nov

Valid

Oleneva

Devonian (EmsianEifelian)

Sharur Formation

 Azerbaijan

A member of Spiriferida belonging to the family Adolfiidae. The type species is G. dagnensis.

Gundaria[68]

Gen. et sp. nov

Valid

Angiolini et al.

Permian

 Tajikistan

The type species is G. insolita.

Harperoides[69]

Gen. et sp. nov

Valid

Baranov & Blodgett

Devonian (Pragian)

Soda Creek Limestone

 United States
( Alaska)

A member of Strophomenida belonging to the subfamily Mesodouvillininae. The type species is Harperoides alaskensis.

Hemileurus politus[68]

Sp. nov

Valid

Angiolini et al.

Permian

 Tajikistan

Holcothyris campbelli[52]

Sp. nov

Valid

MacFarlan

Jurassic

 New Zealand

A member of Terebratulida.

Hustedia shumardi[60]

Sp. nov

Valid

Torres-Martínez, Sour-Tovar & Barragán

Permian (Leonardian)

Paso Hondo Formation

 Mexico

Inflatia coodzavuii[53]

Sp. nov

Valid

Torres-Martínez & Sour-Tovar

Carboniferous (late Mississippian, middle Pennsylvanian)

Ixtaltepec Formation

 Mexico

A member of Productoidea.

Iridistrophia (Flabellistrophia)[70]

Subgen. et comb. sp. nov

Valid

Jansen

Devonian (late Emsian to Eifelian)

 Germany
 Venezuela?

A member of Chilidiopsidae; a subgenus of Iridistrophia. The type species is "Orthis" hipponyx Schnur (1851); the subgenus also includes new species Iridistrophia (Flabellistrophia) musculosa and possibly also "Orthis" undifera Schnur (1853) and Iridistrophia dendritica Benedetto (1984).

Ishimia inflata[49]

Sp. nov

Valid

Popov, Kebriaee-Zadeh & Pour

Ordovician (Darriwilian)

Lashkarak Formation

 Iran

A member of Strophomenida belonging to the family Leptellinidae.

Isogramma nakamurai[63]

Sp. nov

Valid

Tazawa

Permian (Wordian)

 Japan

Jakutoproductus lenensis[71]

Sp. nov

Valid

Makoshin

Early Permian

 Russia

Jakutoproductus talchanensis[71]

Sp. nov

Valid

Makoshin

Early Permian

 Russia

Kjaerina (Kjaerina) gondwanensis[72]

Sp. nov

Valid

Colmenar

Ordovician (late SandbianKatian)

Gabian Formation
Glauzy Formation
Louredo Formation
Portixeddu Formation

 France
 Italy
 Portugal

A rafinesquinid strophomenid brachiopod, a species of Kjaerina.

Kjaerina (Villasina)[72]

Subgen. et 3 sp. et comb. nov

Valid

Colmenar

Ordovician (Katian)

Cavá Formation
Gabian Formation
Portilla de Luna Limestones
Portixeddu Formation
Porto de Santa Anna Formation
Punta Serpeddi Formation
Rosan Formation

 France
 Italy
 Portugal
 Spain

A rafinesquinid strophomenid brachiopod, a subgenus of Kjaerina. The type species of the subgenus is Kjaerina (Villasina) pedronaensis; the subgenus also contains "Hedstroemina" almadenensis Villas (1995), as well as new species Kjaerina (Villasina) meloui and Kjaerina (Villasina) pyrenaica.

Koneviella? fuscina[54]

Sp. nov

Valid

Percival, Engelbretsen & Peng

Cambrian

Huaqiao Formation

 China

A lingulate brachiopod belonging to the family Zhanatellidae.

Kutchithyris challinori[52]

Sp. nov

Valid

MacFarlan

Jurassic

 New Zealand

A member of Terebratulida.

Kutchithyris waitomoensis[52]

Sp. nov

Valid

MacFarlan

Jurassic

 New Zealand

A member of Terebratulida.

Lacunites jaroslavi[73]

Sp. nov

Valid

Mergl & Kraft

Early Ordovician

Klabava Formation

 Czech Republic

A paterinate brachiopod.

Lampazarorthis[64]

Gen. et comb. et sp. nov

Valid[65]

Benedetto & Muñoz

Early Ordovician

 Argentina

A plectorthoid brachiopod. A new genus for "Eoorthis" bifurcata Harrington (1937); genus also includes "Nanorthis" brachymyaria Benedetto in Benedetto & Carrasco (2002), as well as new species Lampazarorthis alata.

Lepidomena multiplicata[49]

Sp. nov

Valid

Popov, Kebriaee-Zadeh & Pour

Ordovician (Darriwilian)

Lashkarak Formation

 Iran

A member of Strophomenida belonging to the family Leptellinidae.

Leptathyris gornensis[50]

Sp. nov

Valid

Baliński in Baliński, Racki & Halamski

Devonian (Frasnian)

 Poland

A member of Athyrididae.

Liaous[74]

Gen. et sp. nov

Valid

He & Chen in He et al.

Middle Triassic (early Anisian)

Xinyuan Formation

 China

A relative of Mentzelia and Paramentzelia. The type species is Liaous shaiwensis.

Loboidothyris awakinoensis[52]

Sp. nov

Valid

MacFarlan

Jurassic

 New Zealand

A member of Terebratulida.

Loboidothyris grantmackiei[52]

Sp. nov

Valid

MacFarlan

Jurassic

 New Zealand

A member of Terebratulida.

Loboidothyris marokopaensis[52]

Sp. nov

Valid

MacFarlan

Jurassic

 New Zealand

A member of Terebratulida.

Lyonia rochacamposi[75]

Sp. nov

Valid

Taboada et al.

Early Permian (latest Asselian-earliest Sakmarian)

Taciba Formation

 Brazil

A member of Productida belonging to the family Auriculispinidae, a species of Lyonia.

Mendozotreta[76]

Gen. et comb. nov

Valid

Holmer et al.

Ordovician

Antelope Valley Limestone
Lindero Formation

 Argentina
 United States
( Nevada)

A member of Acrotretida belonging to the family Acrotretidae. The type species is "Conotreta" devota Krause & Rowell (1975).

Mesoleptostrophia belli[77]

Sp. nov

Valid

Earp

Early Devonian

Montys Hut Formation

 Australia

Minutella bulgarica[78]

Sp. nov

Valid

Bitner & Motchurova-Dekova

Miocene (Badenian)

 Bulgaria

Monelasmina montisjosephi[50]

Sp. nov

Valid

Baliński in Baliński, Racki & Halamski

Devonian (Frasnian)

 Poland

A member of Draboviidae.

Nasakia[79]

Gen. et sp. nov

Valid

Streng et al.

Cambrian

Henson Gletscher Formation

 Greenland

A member of Rhynchonelliformea belonging to the class Obolellata and the order Naukatida. The type species is Nasakia thulensis.

Numericoma rowelli[76]

Sp. nov

Valid

Holmer et al.

Ordovician (Darriwilian)

Antelope Valley Limestone
Ponon Trehue Formation

 Argentina
 United States
( Nevada)

A member of Lingulata belonging to the family Ephippelasmatidae.

Nushbiella kleithria[47]

Sp. nov

Valid

Percival in Percival et al.

Ordovician

 Australia

A member of Lingulata belonging to the family Siphonotretidae.

Obliquorhynchia[80]

Gen. et comb. nov

Valid

Schrøder, Lauridsen & Surlyk

Paleocene (Danian)

Faxe Formation
Vigny Formation

 Denmark
 France
 Sweden

A member of Rhynchonellida belonging to the superfamily Pugnacoidea and the family Basiliolidae; a new genus for "Terebratula" flustracea von Buch (1834).

Orthis dehmollaensis[49]

Sp. nov

Valid

Popov, Kebriaee-Zadeh & Pour

Ordovician (Darriwilian)

Lashkarak Formation

 Iran

A member of Orthida belonging to the family Orthidae.

Paraspirifer (Laurentispirifer)[70]

Subgen. et comb. nov

Valid

Jansen

Middle Devonian

 United States
 Venezuela

A subgenus of Paraspirifer. The type species is Paraspirifer conradi Godefroid & Fagerstrom (1983); the subgenus also includes "Delthyris" acuminata Conrad (1839), "Terebratula" acuminatissima de Castelnau (1843), "Spirifer" bownockeri Stewart (1927), Paraspirifer halli Godefroid & Fagerstrom (1983) and Paraspirifer clarkei Godefroid & Fagerstrom (1983).

Paraspirifer (Mosellospirifer)[70]

Subgen. et comb. nov

Valid

Jansen

Devonian (late Emsian to early Eifelian

 Germany
 Canada?
 China?

A subgenus of Paraspirifer. The type species is Paraspirifer sandbergeri Solle (1971); the subgenus also includes Spirifer auriculatus Sandberger & Sandberger (1856), Paraspirifer sandbergeri longimargo Solle (1971) (elevated to species rank), Paraspirifer eos Solle (1971) and Paraspirifer sandbergeri nepos Solle (1971). The subgenus might also include Paraspirifer gigantea Su (1976) and Paraspirifer desbiensi Bizzarro & Lespérance (1999).

Parazhanatella[54]

Gen. et sp. nov

Valid

Percival, Engelbretsen & Peng

Cambrian

Huaqiao Formation

 China

A lingulate brachiopod belonging to the family Zhanatellidae. The type species is P. paibia.

Pedderia[69]

Gen. et sp. nov

Valid

Baranov & Blodgett

Devonian (Pragian)

Soda Creek Limestone

 United States
( Alaska)

A member of Rhynchonellida belonging to the family Pygmaellidae. The type species is Pedderia fragosa.

Phragmorthis shahrudensis[49]

Sp. nov

Valid

Popov, Kebriaee-Zadeh & Pour

Ordovician (Darriwilian)

Lashkarak Formation

 Iran

A member of Orthida belonging to the family Phragmorthidae.

Prospira pseudostruniana[81]

Sp. nov

Valid

Mottequin & Brice

Devonian (late Famennian)

Etrœungt Formation

 France

A member of Spiriferida belonging to the family Spiriferidae.

Psygmakantha[47]

Gen. et sp. nov

Valid

Percival in Percival et al.

Ordovician

 Australia

A member of Lingulata belonging to the family Ephippelasmatidae. The type species is P. malachiensis'.

Punctospirifer gerankalasus[58]

Sp. nov

Valid

Oleneva

Carboniferous (Tournaisian)

 Azerbaijan

A member of Spiriferinida belonging to the family Punctospiriferidae.

Qaidamospirifer[67]

Gen. et sp. nov

Valid

Chen, Lee & Shi in Shi et al.

Carboniferous (Serpukhovian)

Huaitoulata Formation

 China

A member of Spiriferoidea belonging to the family Choristitidae. The type species is Q. elongatus.

Rafinesquina (Mesogeina)[72]

Subgen. et comb. et 2 sp. nov

Valid

Colmenar

Ordovician (Katian)

Bohdalec Formation
Fombuena Formation
Gabian Formation
Lower Ktaoua Formation
Porto de Santa Anna Formation
Upper Tiouririne Formation
Zahorany Formation

 Czech Republic
 France
 Germany
 Morocco
 Portugal
 Spain

A rafinesquinid strophomenid brachiopod, a subgenus of Rafinesquina. The type species of the subgenus is "Leptaena" pseudoloricata Barrande (1848); the subgenus also includes Rafinesquina pomoides Havlíček (1971), as well as new species Rafinesquina (Mesogeina) gabianensis and Rafinesquina (Mesogeina) loredensis.

Rhipidomella magna[63]

Sp. nov

Valid

Tazawa

Permian (Wordian)

 Japan

Rogorthis? oriens[49]

Sp. nov

Valid

Popov, Kebriaee-Zadeh & Pour

Ordovician (Darriwilian)

Lashkarak Formation

 Iran

A member of Orthida belonging to the family Orthidae.

Sampo suduvensis[59]

Sp. nov

Valid

Paškevičius & Hints

Late Ordovician

 Lithuania
 Russia
( Kaliningrad Oblast)

A member of Strophomenida belonging to the family Leptestiidae.

Sartenaerirhynchus[70]

Gen. et comb. nov

Valid

Jansen

Devonian (middle Siegenian to late Emsian)

 Belgium
 Germany
 Luxembourg

A member of Rhynchonellida belonging to the superfamily Uncinuloidea. The type species is "Terebratula" antiqua Schnur (1853); genus also includes "Uncinulus" frontecostatus Drevermann (1902)

Scaphelasma quadratum[47]

Sp. nov

Valid

Engelbretsen in Percival et al.

Ordovician

 Australia

A member of Lingulata belonging to the family Scaphelasmatidae.

Schizocrania equestra[82]

Sp. nov

Valid

Mergl & Nolčová

Ordovician (Katian)

Bohdalec Formation

 Czech Republic

A member of Discinoidea belonging to the family Trematidae.

Septaliphoria felberi[51]

Sp. nov

Valid

Sulser

Middle Jurassic (Callovian)

  Switzerland

A member of Rhynchonellida belonging to the superfamily Hemithiridoidea and the family Cyclothyrididae.

Siberioproductus[56]

Gen. et sp. nov

Valid

Baranov, Sokiran & Blodgett

Devonian (Famennian)

Chekursk Formation

 Russia
( Sakha Republic)

A member of Rhynchonelliformea belonging to the group Productida and the family Productellidae. The type species is S. boreus.

Skenidioides cretus[50]

Sp. nov

Valid

Halamski in Baliński, Racki & Halamski

Devonian (Frasnian)

 Poland

A member of Skenidiidae.

Spinobolus[83]

Gen. et sp. nov

Valid

Zhang et al.

Cambrian

Shuijingtuo Formation

 China

A linguloid brachiopod. The type species is Spinobolus popovi.

Tapongaspirifer[77]

Gen. et sp. nov

Valid

Earp

Early Devonian

Montys Hut Formation

 Australia

A member of Spiriferida belonging to the family Filispiriferidae. The type species is Tapongaspirifer melodiae.

Taungurungetes[77]

Gen. et comb. nov

Valid

Earp

Early Devonian

Montys Hut Formation
Norton Gully Sandstone

 Australia

A possible member of Devonochonetinae. The type species is "Chonetes" taggertyensis Gill (1945) from the Montys Hut Formation; genus also contains second, unnamed species from the Norton Gully Sandstone.

Terebratulina leeae[52]

Sp. nov

Valid

MacFarlan

Jurassic

 New Zealand

A member of Terebratulida.

Terebratulina putiensis[52]

Sp. nov

Valid

MacFarlan

Jurassic

 New Zealand

A member of Terebratulida.

Thaerodonta notabile[84]

Sp. nov

Valid

Paškevičius

Ordovician (Katian)

Kaimynai Formation

 Lithuania

Thulatrypa[85]

Gen. et sp. et comb. nov

Valid

Huang et al.

Silurian

 China
 Norway

An atrypoid brachiopod. The type species is Thulatrypa gregaria; genus also contains "Meifodia" orientalis Rong, Xu & Yang (1974).

Tomteluva[79]

Gen. et sp. nov

Valid

Streng et al.

Cambrian

Stephen Formation

 Canada
( British Columbia)

A member of Rhynchonelliformea belonging to the class Obolellata and the order Naukatida. The type species is Tomteluva perturbata.

Trentingula[86]

Gen. et 3 sp. et comb. nov

Valid

Posenato

Late Permian and Early Triassic

 Hungary
 Italy
 United States
( Wyoming)

A member of Linguloidea belonging to the family Lingulidae. The type species is T. lorigae; genus also includes new species T. mazzinensis and T. prinothi, as well as Trentingula borealis (Bittner, 1899).

Triangulospirifer[67]

Nom. nov

Valid

Lee in Shi et al.

Carboniferous

 United Kingdom

A member of Spiriferida. A replacement name for Triangularia Poletaev (2001).

Weberproductus[53]

Gen. et sp. nov

Valid

Torres-Martínez & Sour-Tovar

Carboniferous (middle Pennsylvanian)

Ixtaltepec Formation

 Mexico

A member of Productoidea. Genus includes new species W. donajiae.

Wenjukovispirifer[58]

Gen. et comb. nov

Valid

Oleneva

Devonian (Famennian)

 Russia

A member of Spiriferida belonging to the family Cyrtospiriferidae. The type species is "Spirifer" koscharicus Ljaschenko (1959); genus also includes "Spirifer" archiaci Murchison (1840), "Spirifer" brodi Wenjukov (1886) and W. lebedjanicus (Nalivkin, 1947).

Zeilleria opuatiaensis[52]

Sp. nov

Valid

MacFarlan

Jurassic

 New Zealand

A member of Terebratulida.

Zeilleria waiohipaensis[52]

Sp. nov

Valid

MacFarlan

Jurassic

 New Zealand

A member of Terebratulida.

Molluscs

Echinoderms

Name Novelty Status Authors Age Unit Location Notes Images

Alternocidaris[87]

Gen. et comb. nov

Valid

El Qot, Abdelhamid, & Abdelghany

Late Cretaceous (Cenomanian, possibly Turonian)

 Egypt

A sea urchin belonging to the group Camarodonta and the family Glyphocyphidae. The type species is "Tiaridia" weldoni Fourtau (1921).

Applinocrinus russelli[88]

Sp. nov

Valid[89]

Gale

Late Cretaceous (Maastrichtian)

Prairie Bluff Formation

 United States
( Mississippi)

A crinoid belonging to the group Articulata and the family Saccocomidae.

Apsidocrinus doreckae[90]

Sp. nov

Valid

Konieczyński, Pisera & Fózy

Early Cretaceous

 Hungary

A cyrtocrinid crinoid, a species of Apsidocrinus.

Arabicodiadema[91]

Gen. et comb. et sp. nov

Valid

Abdelhamid, El Qot & Abdelghany

Cretaceous (Albian to Cenomanian)

 Oman
 United Arab Emirates

A heterodiadematid sea urchin. The type species is "Trochodiadema" dhofarense Roman (1991); genus also includes new species Arabicodiadema alii.

Atalopegaster[92]

Gen. et sp. nov

Valid

Blake & Guensburg

Late Jurassic (Oxfordian)

Swift Formation

 United States
( Montana)

A starfish belonging to the family Stichasteridae. The type species is A. gundersoni.

Bystrowicrinus (col.) depressus[93]

Sp. nov

Valid

Donovan & Keighley

Late Silurian

West Point Formation

 Canada
( Quebec)

A crinoid.

Calliderma lindneri[94]

Sp. nov

Valid

Niebuhr & Seibertz

Late Cretaceous (Turonian)

Schmilka Formation

 Germany

A starfish belonging to the family Goniasteridae, a species of Calliderma.

Carolinaster[95]

Gen. et sp. nov

Valid

Osborn, Mooi & Ciampaglio

Eocene

Castle Hayne Limestone

 United States
( North Carolina)

A heart urchin belonging to the family Prenasteridae. Genus includes new species C. varnami.

Comptoniaster michaelisi[94][96]

Nom. nov

Valid

Niebuhr & Seibertz

Late Cretaceous (middle Turonian to middle Coniacian)

 Czech Republic
 Germany
 Poland

A starfish belonging to the family Goniasteridae; a replacement name for Asterias schulzii Cotta sensu Roemer (1840, 1841).

Costatocrinus[88]

Gen. et 2 sp. nov

Valid[89]

Gale

Late Cretaceous (Campanian)

Taylor Formation

 United Kingdom
 United States
( Texas)

A crinoid belonging to the group Articulata and the family Saccocomidae. The type species is C. brydonei; genus also includes C. mortimorei.

Cultellacrinus[88]

Gen. et 2 sp. nov

Valid[89]

Gale

Late Cretaceous

 Sweden
 United Kingdom

A crinoid belonging to the group Articulata and the family Roveacrinidae. The type species is C. gladius; genus also includes C. labyrinthus.

Cunidentechinus[97]

Gen. et comb. nov

Valid

Smith

Early Jurassic

 United Kingdom

A stem-carinacean sea urchin; a new genus for "Echinus" minutus Buckman in Murchison (1845).

Darwinaster[98]

Gen. et sp. nov

Valid

Hunter, Rushton & Stone

Early Devonian

Fox Bay Formation

 Falkland Islands

A brittle star related to members of the genus Protaster. Genus includes new species D. coleenbiggsae.

Echinolampas andalusiensis[95]

Sp. nov

Valid

Osborn, Mooi & Ciampaglio

Eocene

Moodys Branch Formation

 United States
( Alabama)

A sea urchin belonging to the family Echinolampadidae.

Gitolampas twitchelli[95]

Sp. nov

Valid

Osborn, Mooi & Ciampaglio

Eocene

Santee Limestone

 United States
( South Carolina)

A sea urchin belonging to the group Cassiduloida.

Glyptocrinus nodosus[99]

Sp. nov

Valid

Kallmeyer & Ausich

Ordovician (Katian)

Kope Formation

 United States
( Kentucky
 Ohio)

A camerate crinoid, a species of Glyptocrinus.

Goniopygus macrotuberculatus[87]

Sp. nov

Valid

El Qot, Abdelhamid, & Abdelghany

Late Cretaceous (Cenomanian)

Raha Formation

 Egypt

A sea urchin belonging to the group Arbacioida and the family Acropeltidae.

Goniopygus subaequalis[87]

Sp. nov

Valid

El Qot, Abdelhamid, & Abdelghany

Late Cretaceous (Cenomanian)

Raha Formation

 Egypt

A sea urchin belonging to the group Arbacioida and the family Acropeltidae.

Haimea brooksi[95]

Sp. nov

Valid

Osborn, Mooi & Ciampaglio

Eocene

Ocala Limestone

 United States
( Florida)

A sea urchin belonging to the group Oligopygoida.

Hessicrinus[88]

Gen. et 2 sp. nov

Valid[89]

Gale

Late Cretaceous (Campanian)

 United Kingdom

A crinoid belonging to the group Articulata and the family Roveacrinidae. The type species is H. filigree; genus also includes H. scalaensis.

Hiiumaacrinus[100]

Gen. et sp. nov

Valid

Ausich & Wilson

Silurian (Rhuddanian)

Hilliste Formation

 Estonia

A crinoid. Genus includes new species H. vinni.

Jakeocrinus[88]

Gen. et sp. nov

Valid[89]

Gale

Late Cretaceous (early Campanian)

Taylor Formation

 United States
( Texas)

A crinoid belonging to the group Articulata and the family Roveacrinidae. The type species is J. ellisensis.

Lakotacrinus[101]

Gen. et sp. nov

Valid

Hunter et al.

Late Cretaceous (Campanian)

Pierre Shale

 United States
( South Dakota)

A stalked crinoid. The type species is L. brezinai.

Lehmannaster[102]

Gen. et sp. nov

Valid

Blake, Guensburg & Lefebvre

Ordovician (late Darriwilian)

Traveusot Formation

 France

A stenuroid asterozoan. Genus includes new species L. spinosus.

Lopidiaster[102]

Gen. et sp. nov

Valid

Blake, Guensburg & Lefebvre

Ordovician (Floian)

Garden City Formation

 United States
( Utah)

A stenuroid asterozoan. Genus includes new species L. jamisoni.

Loriolidiadema[91]

Gen. et comb. nov

Valid

Abdelhamid, El Qot & Abdelghany

Cretaceous (Aptian-Cenomanian)

 Egypt
 Portugal
Border region between  Syria and  Lebanon

A heterodiadematid sea urchin. The type species is "Pseudodiadema" libanoticum de Loriol (1887); genus also includes Loriolidiadema sculptile (de Loriol, 1887).

Lucernacrinus[88]

Gen. et sp. nov

Valid[89]

Gale

Late Cretaceous

 United Kingdom

A crinoid belonging to the group Articulata and the family Roveacrinidae. The type species is L. woodi.

Maennilocystis[103]

Gen. et sp. nov

Valid

Paul & Rozhnov

Late Ordovician

 Estonia

A cystoid belonging to the family Callocystitidae. The type species is M. heckeri.

Manfredaster praebulbiferus[94]

Sp. nov

Valid

Niebuhr & Seibertz

Late Cretaceous (Cenomanian to Coniacian)

Dölzschen Formation
Schrammstein Formation
Strehlen Formation

 Germany

A starfish belonging to the group Valvatida and the family Stauranderasteridae.

Mariania comaschicariae[104]

Sp. nov

Valid

Stara, Borghi & Kroh

Miocene (Aquitanian to early Burdigalian)

Nurallao Formation

 Italy

A heart urchin belonging to the superfamily Spatangoidea, a species of Mariania.

Mariania stefaninii[104]

Sp. nov

Valid

Stara, Borghi & Kroh

Miocene (late Burdigalian to early Langhian)

Pantano Formation

 Italy

A heart urchin belonging to the superfamily Spatangoidea, a species of Mariania.

Meturaster[102]

Gen. et sp. nov

Valid

Blake, Guensburg & Lefebvre

Ordovician (middle Darriwilian)

Lehman Formation

 United States
( Nevada)

A starfish. Genus includes new species M. belli.

Monophoraster telfordi[105]

Sp. nov

Valid

Mooi, Martínez & del Río

Early Miocene

Chenque Formation

 Argentina

A sand dollar.

Nucleolites usoi[106]

Sp. nov

Valid

Forner i Valls

Early Cretaceous (Barremian)

Artoles Formation

 Spain

A sea urchin.

Osteocrinus sinensis[107]

Sp. nov

Valid

Hess, Etter & Hagdorn

Late Triassic (early Carnian)

Xiaowa Formation

 China

A crinoid belonging to the group Roveacrinida, a species of Osteocrinus.

Pentamerocrinus kosovi[108]

Sp. nov

Valid

Rozhnov

Ordovician

 Russia
( Leningrad Oblast)

A crinoid belonging to the group Inadunata, a species of Pentamerocrinus.

Petraster crozonensis[102]

Sp. nov

Valid

Blake, Guensburg & Lefebvre

Ordovician (early Katian)

Kermeur Formation

 France

A starfish.

Phyllocrinus bersekensis[90]

Sp. nov

Valid

Konieczyński, Pisera & Fózy

Early Cretaceous

 Hungary

A cyrtocrinid crinoid, a species of Phyllocrinus.

Placentinechinus[109]

Gen. et sp. nov

Valid[110]

Borghi & Garilli

Pleistocene (GelasianCalabrian)

 Italy

A sea urchin belonging to the family Temnopleuridae. The type species is P. davolii.

Plagiochasma saurai[106]

Sp. nov

Valid

Forner i Valls

Early Cretaceous (Barremian)

Artoles Formation

 Spain

A sea urchin.

Platelicrinus longispinus[88]

Sp. nov

Valid[89]

Gale

Late Cretaceous

 United Kingdom

A crinoid belonging to the group Articulata and the family Roveacrinidae.

Protaeropsis[111]

Nom. nov

Valid

Doweld

Paleocene (Thanetian)

 Spain

A heart urchin; a replacement name for Sphenaster Jeffery in Smith et al. (1999).

Protaxocrinus estoniensis[100]

Sp. nov

Valid

Ausich & Wilson

Silurian (Rhuddanian)

Hilliste Formation

 Estonia

A crinoid.

Pseudomilnia[97]

Gen. et comb. nov

Valid

Smith

Middle Jurassic

 France
 United Kingdom

A salenioidan sea urchin; a new genus for "Acrosalenia" miliaria Paris (1908).

Sagittacrinus[88]

Gen. et sp. nov

Valid[89]

Gale

Late Cretaceous (early Campanian)

Taylor Formation

 United Kingdom
 United States
( Texas)

A crinoid belonging to the group Articulata and the family Saccocomidae. The type species is S. torpedo.

Salenia buddyi[95]

Sp. nov

Valid

Osborn, Mooi & Ciampaglio

Eocene

Santee Limestone

 United States
( South Carolina)

A sea urchin belonging to the group Salenioida.

Schuchertia sprinklei[102]

Sp. nov

Valid

Blake, Guensburg & Lefebvre

Ordovician (Dapingian)

Kanosh Shale

 United States
( Utah)

A starfish.

Stellacrinus[88]

Gen. et 2 sp. nov

Valid[89]

Gale

Late Cretaceous (Santonian and Campanian)

Gulpen Formation
Taylor Formation
Vaals Formation

 Netherlands
 Sweden
 United Kingdom
 United States
( Texas)

A crinoid belonging to the group Articulata and the family Roveacrinidae. The type species is S. hugesae; genus also includes S. pannosus.

Stomechinus phillipsii[97]

Sp. nov

Valid

Smith

Jurassic

 United Kingdom

A sea urchin.

Thinocrinus akanthos[112]

Sp. nov

Valid

Rhenberg, Ausich & Meyer

Carboniferous (Mississippian)

Fort Payne Formation

 United States
( Kentucky)

A camerate crinoid belonging to the group Monobathrida and the family Actinocrinitidae.

Trochotiara aalensis[97]

Sp. nov

Valid

Smith

Middle Jurassic

 United Kingdom

A sea urchin.

Wrightechnus[97]

Gen. et comb. nov

Valid

Smith

Middle Jurassic

 France
 United Kingdom

A sea urchin, possibly a stem-salenioidan; a new genus for "Acrosalenia" lycetti Wright (1851).

Conodonts

Name Novelty Status Authors Age Unit Location Notes Images

Amydrotaxis corniculans truncus[113]

Subsp. nov

Valid

Mathieson et al.

Early Devonian

Mountain Dam Limestone

 Australia

A member of Ozarkodinida belonging to the family Spathognathodontidae.

Cypricriodus[114][115]

Gen. et comb. nov

Valid

Murphy, Carls & Valenzuela-Ríos

Silurian and Early Devonian

 Australia
 United States
( Nevada)

Genus includes "Caudicriodus" hesperius (Klapper & Murphy, 1975).

Eognathodus kuangi[116]

Sp. nov

Valid

Earp

Early Devonian

 China

Eognathodus sulcatus lanei[113]

Subsp. nov

Valid

Mathieson et al.

Early Devonian

Mountain Dam Limestone

 Australia

A member of Ozarkodinida belonging to the family Spathognathodontidae.

Eognathodus sulcatus sicatus[113]

Subsp. nov

Valid

Mathieson et al.

Early Devonian

Booth Limestone

 Australia

A member of Ozarkodinida belonging to the family Spathognathodontidae.

Gladigondolella okayi[117]

Sp. nov

Valid

Kiliç

Triassic

Kayabaşı Limestone

 Turkey

Gondolella wardlawi[118]

Sp. nov

Valid

Nestell & Pope in Nestell, Wardlaw & Pope

Carboniferous (Pennsylvanian)

Excello Shale

 United States
( Iowa)

Heliagnathus[113]

Gen. et sp. nov

Valid

Mathieson et al.

Early Devonian

Mountain Dam Limestone

 Australia

A member of Ozarkodinida of uncertain phylogenetic placement. The type species is H. parvilabiatus.

Icriodus ampliatus[113]

Sp. nov

Valid

Mathieson et al.

Early Devonian

Mountain Dam Limestone

 Australia

A member of Prioniodontida belonging to the family Icriodontidae.

Icriodus michiganus[119]

Sp. nov

Valid

Narkiewicz & Bultynck

Devonian (Eifelian)

Michigan Basin

 United States

Icriodus praealternatus ferus[120]

Subsp. nov

Valid

Wang et al.

Late Devonian (Frasnian/Famennian boundary)

Hongguleleng Formation

 China

A subspecies of Icriodus praealternatus.

Icriodus plurinodosus[120]

Sp. nov

Valid

Wang et al.

Late Devonian (early Famennian)

Hongguleleng Formation

 China

A species of Icriodus.

Icriodus stenoancylus junggarensis[120]

Subsp. nov

Valid

Wang et al.

Late Devonian (early Famennian)

Hongguleleng Formation

 China

A subspecies of Icriodus stenoancylus.

Idiognathodus lateralis[121]

Sp. nov

Valid

Hogancamp, Barrick & Strauss

Carboniferous (early Gzhelian)

 China
 Russia
 Ukraine
 United States

A member of Ozarkodinida belonging to the family Idiognathodontidae, a species of Idiognathodus.

Idiognathodus tuberis[118]

Sp. nov

Valid

Nestell, Wardlaw & Pope

Carboniferous (Pennsylvanian)

Excello Shale

 United States
( Iowa)

Neogondolella cuneiforme[122]

Sp. nov

Valid

Golding & Orchard

Middle Triassic (Anisian)

 Canada
( British Columbia)

Neogondolella curva[122]

Sp. nov

Valid

Golding & Orchard

Middle Triassic (Anisian)

 Canada
( British Columbia)
 United States
( Nevada)

Neogondolella dilacerata[122]

Sp. nov

Valid

Golding & Orchard

Middle Triassic (Anisian)

 Canada
( British Columbia)

Originally described as a species of Neogondolella, but subsequently transferred to the genus Magnigondolella.[123]

Neogondolella hastata[122]

Sp. nov

Valid

Golding & Orchard

Middle Triassic (Anisian)

 Canada
( British Columbia)
 United States
( Nevada)

Neogondolella indicta[122]

Sp. nov

Valid

Golding & Orchard

Middle Triassic (Anisian)

 Canada
( British Columbia)
 United States
( Nevada)

Neogondolella panlaurentia[122]

Sp. nov

Valid

Golding & Orchard

Middle Triassic (Anisian)

 Canada
( British Columbia)
 United States
( Nevada)

Neogondolella tenera[122]

Sp. nov

Valid

Golding & Orchard

Middle Triassic (Anisian)

 Canada
( British Columbia)
 United States
( Nevada)

Neogondolella vellicata[122]

Sp. nov

Valid

Golding & Orchard

Middle Triassic (Anisian)

 Canada
( British Columbia)
 United States
( Nevada)

Neopolygnathus huijunae[120]

Sp. nov

Valid

Wang et al.

Late Devonian (early Famennian)

Hongguleleng Formation

 China

A species of Neopolygnathus.

Neospathodus planus[124]

Sp. nov

Valid

Chen et al.

Early Triassic

 Slovenia

Oulodus astriatus[113]

Sp. nov

Valid

Mathieson et al.

Early Devonian

Booth Limestone

 Australia

A member of Prioniodinida belonging to the family Prioniodinidae.

Ozarkodina multistriola[113]

Sp. nov

Valid

Mathieson et al.

Early Devonian

Mountain Dam Limestone

 Australia

A member of Ozarkodinida belonging to the family Spathognathodontidae.

Ozarkodina scoliciformis[113]

Sp. nov

Valid

Mathieson et al.

Early Devonian

Mountain Dam Limestone

 Australia

A member of Ozarkodinida belonging to the family Spathognathodontidae.

Ozarkodina selfi cordata[113]

Subsp. nov

Valid

Mathieson et al.

Early Devonian

Mountain Dam Limestone

 Australia

A member of Ozarkodinida belonging to the family Spathognathodontidae.

Ozarkodina selfi lanceola[113]

Subsp. nov

Valid

Mathieson et al.

Early Devonian

Mountain Dam Limestone

 Australia

A member of Ozarkodinida belonging to the family Spathognathodontidae.

Ozarkodina selfi lenticula[113]

Subsp. nov

Valid

Mathieson et al.

Early Devonian

Mountain Dam Limestone

 Australia

A member of Ozarkodinida belonging to the family Spathognathodontidae.

Panderodus rhytiodus[113]

Sp. nov

Valid

Mathieson et al.

Early Devonian

Mountain Dam Limestone

 Australia

A member of the family Panderodontidae.

Pelekysgnathus inequalis[113]

Sp. nov

Valid

Mathieson et al.

Early Devonian

Booth Limestone

 Australia

A member of Prioniodontida belonging to the family Icriodontidae.

Pelekysgnathus jeppssoni[125]

Sp. nov

Valid

Nazarova

Devonian (Eifelian)

 Russia
( Bryansk Oblast)

Platyvillosus corniger[124]

Sp. nov

Valid

Chen et al.

Early Triassic

 Slovenia

Polygnathus abaimovae[126]

Sp. nov

Valid

Baranov & Blodgett

Devonian (Emsian)

 Russia

Polygnathus beckeri[126]

Nom. nov

Valid

Baranov & Blodgett

Devonian (Emsian)

 Russia

A replacement name for Polygnathus inflexus Baranov (1992).

Polygnathus datnensis[126]

Sp. nov

Valid

Baranov & Blodgett

Devonian (Emsian)

 Russia

Polygnathus dogdensis[126]

Sp. nov

Valid

Baranov & Blodgett

Devonian (Emsian)

 Russia

Polygnathus novozemelicus[126]

Sp. nov

Valid

Baranov & Blodgett

Devonian (Emsian)

 Russia

“Polygnathus” pseudocommunis[120]

Sp. nov

Valid

Wang et al.

Late Devonian (early Famennian)

Hongguleleng Formation

 China

Possibly a species of Polygnathus.

Polygnathus slastenovi[126]

Sp. nov

Valid

Baranov & Blodgett

Devonian (Emsian)

 Russia

Polygnathus tarabukini[126]

Sp. nov

Valid

Baranov & Blodgett

Devonian (Emsian)

 Russia

Siphonodella (Siphonodella) jii[127]

Nom. nov

Disputed

Becker, Kaiser & Aretz

Carboniferous (Tournaisian)

Chappel Limestone

 United States

A replacement name for Siphonodella (Siphonodella) hassi Ji (1985). Plotitsyn and Zhuravlev (2016) considered S. jii to be a junior synonym of Siphonodella quadruplicata (Branson et Mehl), because they considered the specimen selected as the holotype of S. jii to be more likely to represent an ontogenetic stage of S. quadruplicata.[128]

Staeschegnathus[129]

Gen. et sp. nov

Valid

Koike

Triassic

Taho Formation

 Japan

An ellisonid conodont. The type species is S. perrii.

Fishes

Amphibians

Research

New taxa

Temnospondyls[edit]

Name Novelty Status Authors Age Unit Location Notes Images

Cyclotosaurus buechneri[139]

Sp. nov

Valid

Witzmann, Sachs & Nyhuis

Late Triassic (middle Carnian)

Stuttgart Formation

 Germany

A mastodonsauroid temnospondyl, a species of Cyclotosaurus.

Konzhukovia sangabrielensis[140]

Sp. nov

Valid[141]

Pacheco et al.

Permian (early Guadalupian)

Rio do Rasto Formation

 Brazil

Samarabatrachus[142]

Gen. et sp. nov

Valid

Novikov

Early Triassic

Sukhorechka Formation

 Russia
( Samara Oblast)

A capitosaurid temnospondyl. The type species is S. bjerringi.

Selenocara rossica[142]

Sp. nov

Valid

Novikov

Early Triassic

Sukhorechka Formation

 Russia
( Orenburg Oblast
 Samara Oblast)

A capitosaurid temnospondyl.

Stanocephalosaurus amenasensis[143]

Sp. nov

Valid

Dahoumane et al.

Early-Middle Triassic

Zarzaïtine Series, Illizi Basin

 Algeria

A mastodonsauroid temnospondyl, a species of Stanocephalosaurus.

Syrtosuchus[142]

Gen. et comb. et sp. nov

Valid

Novikov

Early Triassic

Sukhorechka Formation

 Russia
( Orenburg Oblast
 Samara Oblast)

A benthosuchid temnospondyl. The type species is "Wetlugasaurus" samarensis Sennikov (1981); genus also includes new species S. morkovini.

Tomeia[144]

Gen. et sp. nov

Valid

Eltink, Da-Rosa & Dias-da-Silva

Early Triassic

Sanga do Cabral Supersequence

 Brazil

A mastodonsauroid temnospondyl. The type species is T. witecki.

Yuanansuchus maopingchangensis[145]

Sp. nov

Valid

Liu

Middle Triassic (Anisian)

Badong Formation

 China

A mastodonsauroid temnospondyl, a species of Yuanansuchus.

Lissamphibians[edit]

Name Novelty Status Authors Age Unit Location Notes Images

Hyogobatrachus[146]

Gen. et sp. nov

Valid

Ikeda, Ota & Matsui

Early Cretaceous

Sasayama Group

 Japan

A frog. The type species is Hyogobatrachus wadai.

Kiyatriton krasnolutskii[147]

Sp. nov

Valid

Skutschas

Middle Jurassic (Bathonian)

Itat Formation

 Russia
( Krasnoyarsk Krai)

A salamander, a species of Kiyatriton.

Kuruleufenia[148]

Gen. et sp. nov

Valid

Gómez

Late Cretaceous (late Campanian–early Maastrichtian)

Allen Formation

 Argentina

A member of Pipidae. The type species is Kuruleufenia xenopoides.

Litoria lundeliusi[149]

Sp. nov

Valid

Tyler & Prideaux

Pleistocene

 Australia

A species of Litoria.

Nuominerpeton[150]

Gen. et sp. nov

Valid

Jia & Gao

Early Cretaceous (BarremianAptian)

Guanghua Formation

 China

A cryptobranchoid salamander of uncertain phylogenetic placement. The type species is N. aquilonaris.

Palaeobatrachus eurydices[151]

Sp. nov

Valid

Villa et al.

Early Pleistocene (Gelasian)

 Netherlands

Phosphotriton[152]

Gen. et sp. nov

Valid

Tissier, Rage, Boistel, Fernandez, Pollet, Garcia and Laurin

Eocene

 France

A salamander. The type species is Phosphotriton sigei.

Prospea[153]

Gen. et sp. nov

Valid

Chen et al.

Paleocene

Naranbulak Formation

 Mongolia

A member of Scaphiopodidae. The type species is Prospea holoserisca.

Qinglongtriton[154]

Gen. et sp. nov

Valid

Jia & Gao

Late Jurassic (Oxfordian)

Tiaojishan Formation

 China

A basal member of Salamandroidea. The type species is Qinglongtriton gangouensis.

Tambabatrachus[146]

Gen. et sp. nov

Valid

Ikeda, Ota & Matsui

Early Cretaceous

Sasayama Group

 Japan

A frog. The type species is Tambabatrachus kawazu.

Others[edit]

Name Novelty Status Authors Age Unit Location Notes Images

Aytonerpeton[155]

Gen. et sp. nov

Valid

Otoo, Clack & Smithson in Clack et al.

Carboniferous (Tournaisian)

Ballagan Formation

 United Kingdom

An early tetrapod of uncertain phylogenetic placement. The type species is A. microps.

Diploradus[155]

Gen. et sp. nov

Valid

Clack & Smithson in Clack et al.

Carboniferous (Tournaisian)

Ballagan Formation

 United Kingdom

An early tetrapod of uncertain phylogenetic placement. The type species is D. austiumensis.

Koilops[155]

Gen. et sp. nov

Valid

Clack & Smithson in Clack et al.

Carboniferous (Tournaisian)

Ballagan Formation

 United Kingdom

An early tetrapod of uncertain phylogenetic placement. The type species is K. herma.

Ossirarus[155]

Gen. et sp. nov

Valid

Clack & Smithson in Clack et al.

Carboniferous (Tournaisian)

Ballagan Formation

 United Kingdom

An early tetrapod of uncertain phylogenetic placement. The type species is O. kierani.

Perittodus[155]

Gen. et sp. nov

Valid

Clack & Smithson in Clack et al.

Carboniferous (Tournaisian)

Ballagan Formation

 United Kingdom

An early tetrapod of uncertain phylogenetic placement. The type species is P. apsconditus.

Lepidosaurs

Lizards

Research[edit]

  • Twelve specimens of lizards (including stem-gekkotans, crown-agamids, a lacertoid, a stem-chamaeleonid and squamates of uncertain phylogenetic placement, probably stem-squamates) are described from the Cretaceous (Albian-Cenomanian boundary) amber from Myanmar by Daza et al. (2016).[156]
  • A study of almost 30 specimens of Polyglyphanodon sternbergi, including almost complete skeletons, is published by Simões et al. (2016), who report the discovery of previously unrecognized ontogenetic series, sexual dimorphism and a complete lower temporal bar in the skull of members of this species.[157]
  • New anatomical data on the Late Cretaceous lizard Slavoia darevskii is published by Tałanda (2016), who interprets it as a stem-amphisbaenian.[158]
  • A study on the skull anatomy of the Eocene amphisbaenian Spathorhynchus fossorium is published by Müller, Hipsley & Maisano (2016).[159]
  • A redescription of the mosasaur Hainosaurus bernardi Dollo (1885) is published by Jimenez-Huidobro & Caldwell (2016), who transfer this species to the genus Tylosaurus and synonymize genera Tylosaurus and Hainosaurus.[160]
  • A revision of the species assigned to the mosasaur genus Tylosaurus is published by Jiménez-Huidobro, Simões & Caldwell (2016);[161] their conclusion that T. kansasensis is a junior synonym of T. nepaeolicus is subsequently rejected by Stewart & Mallon (2018).[162]
  • Early Miocene chamaeleonid fossils, including a specimen tentatively attributed to the species Chamaeleo cf. andrusovi Čerňanský (2010), previously known only from the early Miocene of the Czech Republic, are described from the Aliveri locality (Euboea, Greece) by Georgalis, Villa & Delfino (2016).[163]
  • Lizard fossils which might be the oldest known chameleon fossils from India are described from the Miocene Nagri Formation by Sankhyan & Čerňanský (2016).[164]

New taxa[edit]

Name Novelty Status Authors Age Unit Location Notes Images

Bagaluus[165]

Gen. et sp. nov

Valid

Alifanov

Early Cretaceous

 Mongolia

A member of Scincomorpha belonging to the family Hodzhakuliidae. The type species is B. primigenius.

Carnoscincus[165]

Gen. et sp. nov

Valid

Alifanov

Early Cretaceous

 Mongolia

A member of Scincomorpha belonging to the family Hodzhakuliidae. The type species is C. eublepharus.

Janosikia[166]

Gen. et comb. nov

Valid

Čerňanský, Klembara & Smith

Early Miocene

 Germany

A member of Lacertidae; a new genus for "Ophisaurus" ulmensis Gerhardt (1903).

Jeddaherdan[167]

Gen. et sp. nov

Apesteguía et al.

Late Cretaceous (Cenomanian)

Kem Kem Beds

 Morocco

An iguanian belonging to the group Acrodonta, possibly a relative of the uromasticine agamids. The type species is J. aleadonta.

Ophisauromimus[168]

Gen. et comb. nov

Valid

Čerňanský, Klembara & Műller

Oligocene

 France
 Germany

A member of Anguidae. A new genus for "Dopasia" coderetensis Augé (2005); genus also includes "Dopasia" frayssensis Augé (2005).

Platynotoides[165]

Gen. et sp. nov

Junior homonym

Alifanov

Early Cretaceous

 Mongolia

A member of Scincomorpha belonging to the family Hodzhakuliidae. The type species is P. altidentatus. The generic name is preoccupied by Platynotoides Kaszab (1975).

Pluridens calabaria[169]

Sp. nov

Valid

Longrich

Late Cretaceous (late Campanian)

Nkporo Shale

 Nigeria

A mosasaur, a species of Pluridens.

Solastella[170]

Gen. et sp. nov

Valid

Stocker & Kirk

Eocene

Devil's Graveyard Formation

 United States
( Texas)

A rhineurid amphisbaenian. The type species is Solastella cookei.

Snakes

Research[edit]

  • Lee et al. (2016) examine the limb anatomy of Tetrapodophis amplectus, which according to the authors is suggestive of aquatic habits.[171]
  • A redescription of the Cenomanian snake Simoliophis rochebrunei on the basis of new fossil material from France is published by Rage, Vullo & Néraudeau (2016).[172]
  • Smith & Scanferla (2016) describe a juvenile specimen of Palaeopython fischeri from the Eocene Messel pit with preserved stomach contents, including a specimen of the stem-basilisk species Geiseltaliellus maarius, which in turn preserves an unidentified insect in its stomach.[173]
  • McNamara et al. (2016) describe pigment cells responsible for coloration and patterning preserved in a fossil skin of a colubrid snake from the Late Miocene Libros Lagerstätte (Teruel, Spain).[174]
  • New fossil material of the viperid Laophis crotaloides is described from Greece by Georgalis et al. (2016).[175]

New taxa[edit]

Name Novelty Status Authors Age Unit Location Notes Images

Lunaophis[176]

Gen. et sp. nov

Valid

Albino, Carrillo-Briceño & Neenan

Late Cretaceous (Cenomanian)

La Luna Formation

 Venezuela

A snake of uncertain phylogenetic placement. The type species is L. aquaticus.

Platyspondylophis[177]

Gen. et sp. nov

Valid

Smith et al.

Eocene (Ypresian)

Cambay Shale Formation

 India

A member of Madtsoiidae. The type species is P. tadkeshwarensis.

Rieppelophis[178]

Gen. et comb. nov

Valid

Scanferla, Smith & Schaal

Eocene

Messel pit

 Germany

A member of Boidae. A new genus for "Messelophis" ermannorum Schaal & Baszio (2004).

Ichthyosauromorphs

Research

  • A study of taxonomic richness, disparity and evolutionary rates of ichthyosaurs throughout the Cretaceous period is published by Fischer et al. (2016).[179]
  • A restudy of "Platypterygius" campylodon is published by Fischer (2016), who transfers this species to the genus Pervushovisaurus.[180]
  • A revision of the ichthyosaur material of the British Middle and Late Jurassic referable to Ophthalmosaurus icenicus is published by Moon & Kirton (2016).[181]

New taxa

Name Novelty Status Authors Age Unit Location Notes Images

Cryopterygius kielanae[182]

Sp. nov

Valid

Tyborowski

Late Jurassic (Tithonian)

Kcynia Formation

 Poland

A member of Ophthalmosauridae. Transferred to the genus Undorosaurus by Zverkov & Efimov (in press).[183]

Ichthyosaurus larkini[184]

Sp. nov

Valid[185]

Lomax & Massare

Early Jurassic (Hettangian)

 United Kingdom

Ichthyosaurus somersetensis[184]

Sp. nov

Valid[185]

Lomax & Massare

Early Jurassic (Hettangian)

 United Kingdom

Sclerocormus[186]

Gen. et sp. nov

Valid

Jiang et al.

Early Triassic (Olenekian)

Nanlinghu Formation

 China

A basal member of Ichthyosauriformes. The type species is S. parviceps.

Wahlisaurus[187]

Gen. et sp. nov

Valid[188]

Lomax

Early Jurassic (Hettangian)

 United Kingdom

A member of Leptonectidae. The type species is W. massarae.

Sauropterygians

Research

New taxa

Name Novelty Status Authors Age Unit Location Notes Images

Alexandronectes[193]

Gen. et sp. nov

Valid

Otero et al.

Late Cretaceous (early Maastrichtian)

Conway Formation

 New Zealand

An aristonectine elasmosaurid plesiosaur. The type species is Alexandronectes zealandiensis.

Dawazisaurus[194]

Gen. et sp. nov

Valid

Cheng et al.

Middle Triassic (Anisian)

Guanling Formation

 China

A non-pistosauroid eosauropterygian of uncertain phylogenetic placement. The type species is Dawazisaurus brevis.

Kawanectes[195]

Gen. et comb. nov

Valid

O'Gorman

Late Cretaceous (late Campanian–early Maastrichtian)

Allen Formation

 Argentina

An elasmosaurid plesiosaur. The type species is "Trinacromerum" lafquenianum Gasparini & Goñi (1985).

Lariosaurus vosseveldensis[196]

Sp. nov

Valid

Klein et al.

Middle Triassic (Anisian)

 Netherlands

Polycotylus sopozkoi[197]

Sp. nov

Valid

Efimov, Meleshin & Nikiforov

Late Cretaceous

 Russia

Stenorhynchosaurus[198]

Gen. et sp. nov

Valid

Páramo et al.

Early Cretaceous (late Barremian)

Paja Formation

 Colombia

A pliosaurid plesiosaur. The type species is Stenorhynchosaurus munozi.

Turtles

Research

New taxa

Name Novelty Status Authors Age Unit Location Notes Images

Algorachelus[207]

Gen. et sp. nov

Valid

Pérez-García[208]

Late Cretaceous (Cenomanian)

Arenas de Utrillas Formation

 Portugal[209]  Spain

A member of Bothremydidae. The type species is A. peregrinus.

Anhuichelys doumuensis[210]

Sp. nov

Valid

Tong et al.

Middle Paleocene

Doumu Formation

 China

A stem-tortoise, a species of Anhuichelys.

Clemmys hutchensorum[211]

Sp. nov

Valid

Bourque

Early Pleistocene (late Blancan)

 United States
( Florida)

A species of Clemmys.

Fontainechelon[212]

Gen. et comb. nov

Valid

Pérez-García, Ortega & Jiménez Fuentes

Early Eocene

 France

A tortoise; a new genus for "Achilemys" cassouleti Claude & Tong (2004).

Inaechelys[213]

Gen. et sp. nov

Disputed

Carvalho, Ghilardi & Barreto

Paleocene (Danian)

Maria Farinha Formation

 Brazil

A member of Bothremydidae. The type species is I. pernambucensis. Its status as a valid taxon was challenged by Romano (2016), who considered the genus Inaechelys to be a junior synonym of the genus Rosasia and the species I. pernambucensis/Rosasia pernambucensis to be a nomen dubium.[214]

Keuperotesta[215]

Gen. et sp. nov

Valid

Szczygielski & Sulej

Late Triassic

 Germany

A member of Proterochersidae. The type species is Keuperotesta limendorsa. The genus Keuperotesta was considered to be a junior synonym of the genus Proterochersis by Joyce (2017), though the author maintained K. limendorsa as a distinct species within the latter genus.[216]

Kinosternon notolophus[217]

Sp. nov

Valid

Bourque

Miocene (Clarendonian)

Alachua Formation
Statenville Formation

 United States
( Florida)

A mud turtle.

Kinosternon pannekollops[217]

Sp. nov

Valid

Bourque

Miocene (Clarendonian)

Ogallala Formation

 United States
( Texas)

A mud turtle.

Kinosternon rincon[217]

Sp. nov

Valid

Bourque

Miocene (late Barstovian)

Cerro Conejo Formation

 United States
( New Mexico)

A mud turtle.

Kinosternon wakeeniense[217]

Sp. nov

Valid

Bourque

Miocene (Clarendonian)

Ash Hollow Formation
Ogallala Formation

 United States
( Kansas,
 Nebraska)

A mud turtle.

Neurankylus notos[218]

Sp. nov

Valid

Lichtig & Lucas

Late Cretaceous (Coniacian-Santonian)

Crevasse Canyon Formation

 United States
( New Mexico)

A member of Baenidae.

Neurankylus torrejonensis[219]

Sp. nov

Valid

Lyson et al.

Paleocene (Torrejonian)

Nacimiento Formation

 United States
( New Mexico)

A member of Baenidae.

Notoemys tlaxiacoensis[220]

Sp. nov

Valid[221]

López-Conde et al.

Late Jurassic (Kimmeridgian)

Sabinal Formation

 Mexico

A member of Platychelyidae.

Paiutemys[222]

Gen. et sp. nov

Disputed

Joyce, Lyson & Kirkland

Late Cretaceous (late Cenomanian)

Naturita Formation

 United States
( Utah)

A member of Bothremydidae. The type species is P. tibert. Pérez-García (2018) considered the genus Paiutemys to be a junior synonym of the genus Algorachelus, and transferred the species P. tibert to the latter genus.[223]

Palaeoamyda[224]

Gen. et comb. nov

Valid

Cadena

Eocene

 Germany

A relative of trionychids; a new genus for "Trionyx" messelianus Reinach (1900).

Pelorochelon[212]

Gen. et sp. et comb. nov

Valid

Pérez-García, Ortega & Jiménez Fuentes

Middle Eocene

 Germany
 Spain

A tortoise. The type species is P. soriana; genus also includes Pelorochelon eocaenica (Hummel, 1935).

Proterochersis porebensis[215]

Sp. nov

Valid

Szczygielski & Sulej

Late Triassic

 Poland

A member of Proterochersidae.

Sichuanchelys palatodentata[225]

Sp. nov

Valid

Joyce et al.

Late Jurassic (Oxfordian)

Shishugou Formation

 China

A basal member of Testudinata.

Tartaruscola[226]

Gen. et sp. nov

Valid

Pérez-García

Eocene (Ypresian)

 France

A member of Bothremydidae belonging to the group Foxemydina. The type species is T. teodorii.

Yelmochelys[227]

Gen. et sp. nov

Valid

Brinkman et al.

Late Cretaceous (late Campanian and early Maastrichtian)

Cañon del Tule Formation
Cerro del Pueblo Formation

 Mexico

A stem-kinosternid. The type species is Yelmochelys rosarioae.

Archosauriformes

Basal archosauriforms

Research[edit]

  • A study on the resting metabolic rate of 14 taxa of fossil archosauromorph reptiles as indicated by bone histology is published by Legendre et al. (2016).[228]
  • A study of the phylogenetic relationships of the archosauriforms traditionally assigned to the family Euparkeriidae is published by Sookias (2016).[229]
  • A redescription of the braincase and the inner ear of Euparkeria capensis is published by Sobral et al. (2016).[230]
  • A study of the phylogenetic relationships of archosauromorph reptiles, with an emphasis on the phylogenetic relationships of proterosuchids and erythrosuchids, is published by Ezcurra (2016).[231]
  • A study on the patterns of morphological diversity of the skulls of late Permian to Early Jurassic archosauromorph reptiles is published by Foth et al. (2016).[232]
  • A study on the braincase anatomy of the type specimens of Pseudochampsa ischigualastensis and Tropidosuchus romeri is published by Trotteyn & Paulina-Carabajal (2016).[233]
  • A reevaluation of the neotype specimen of Parasuchus hislopi and a study of the phylogenetic relationships of the species is published by Kammerer et al. (2016), who consider the genus Parasuchus to be a senior synonym of the genera Paleorhinus and Arganarhinus, and refer the species Paleorhinus bransoni Williston (1904), Francosuchus angustifrons Kuhn (1936) and Paleorhinus magnoculus Dutuit (1977) to the genus Parasuchus.[234]
  • A study on the endocranial anatomy (including the brain, inner ear, neurovascular structures and sinus systems) of Parasuchus angustifrons and Ebrachosuchus neukami is published by Lautenschlager & Butler (2016).[235]

New taxa[edit]

Name Novelty Status Authors Age Unit Location Notes Images

Litorosuchus[236]

Gen. et sp. nov

Valid

Li et al.

Middle Triassic

Falang Formation

 China

Probably a relative of Vancleavea. The type species is L. somnii.

Triopticus[237]

Gen. et sp. nov

Valid

Stocker et al.

Late Triassic (latest Carnian-early Norian)

Dockum Group

 United States
( Texas)

Probably a basal member of Archosauriformes. The type species is T. primus.

Pseudosuchians

Research[edit]

New taxa[edit]

Name Novelty Status Authors Age Unit Location Notes Images

Agaresuchus[262]

Gen. et sp. et comb. nov

Valid

Narváez et al.

Late Cretaceous (late CampanianMaastrichtian)

 Spain

A member of Allodaposuchidae. Genus includes new species Agaresuchus fontisensis, as well as Allodaposuchus” subjuniperus.

Bayomesasuchus[263]

Gen. et sp. nov

Valid

Barrios, Paulina-Carabajal & Bona

Late Cretaceous

Cerro Lisandro Formation

 Argentina

A peirosaurid crocodyliform. The type species is Bayomesasuchus hernandezi.

Elosuchus broinae[264]

Sp. nov

Valid

Meunier & Larsson

Late Cretaceous (Cenomanian)

 Algeria

Fortignathus[265]

Gen. et comb. nov

Valid[266]

Young et al.

Cretaceous (late Albian-early Cenomanian)

Echkar Formation

 Niger

A dyrosaurid or a relative of dyrosaurids; a new genus for "Elosuchus" felixi de Lapparent de Broin (2002).

Gryposuchus pachakamue[267]

Sp. nov

Valid

Salas-Gismondi et al.

Miocene

Pebas Formation

 Peru

A member of Gryposuchinae, a species of Gryposuchus.

Kalthifrons[268]

Gen. et sp. nov

Valid

Yates & Pledge

Pliocene

Tirari Formation

 Australia

A member of Mekosuchinae. The type species is K. aurivellensis.

Kentisuchus astrei[269]

Sp. nov

Valid

Jouve

Eocene (late Lutetian)

 France

A member of Tomistominae, a species of Kentisuchus.

Lavocatchampsa[270]

Gen. et sp. nov

Valid

Martin & De Lapparent De Broin

Cretaceous (Albian-Cenomanian)

Kem Kem Beds

 Morocco

A notosuchian. The type species is L. sigogneaurusselae.

Llanosuchus[271]

Gen. et sp. nov

Valid

Fiorelli et al.

Late Cretaceous (Campanian?)

Los Llanos Formation

 Argentina

A notosuchian crocodyliform. The type species is Llanosuchus tamaensis.

Machimosaurus rex[272]

Sp. nov

Valid

Fanti et al.

Early Cretaceous

 Tunisia

A teleosaurid crocodylomorph, a species of Machimosaurus.

Patagosuchus[273]

Gen. et sp. nov

Valid

Lio et al.

Late Cretaceous (Turonian–Coniacian)

Portezuelo Formation

 Argentina

A peirosaurid crocodylomorph. The type species is Patagosuchus anielensis.

Protoalligator[274]

Gen. et comb. nov

Valid

Wang, Sullivan & Liu

Middle Paleocene

Wanghudun Formation

 China

A member of Alligatoroidea of uncertain phylogenetic placement; a new genus for "Eoalligator" huiningensis Young (1982).

Sabinosuchus[275]

Gen. et sp. nov

Valid

Shiller, Porras-Muzquiz & Lehman

Late Cretaceous (Maastrichtian)

Escondido Formation

 Mexico

A member of Dyrosauridae. The type species is S. coahuilensis.

Sabresuchus[276]

Gen. et comb. nov

Valid

Tennant, Mannion & Upchurch

Cretaceous (late BarremianMaastrichtian)

 Romania
 Spain

A member of Paralligatoridae. The type species is "Theriosuchus" ibericus Brinkmann (1989); genus also includes "Theriosuchus" sympiestodon Martin, Rabi & Csiki (2010).

Scutarx[277][278]

Gen. et sp. nov

Valid

Parker

Late Triassic (middle Norian)

Chinle Formation
Cooper Canyon Formation

 United States
( Arizona,  Texas)

An aetosaur. The type species is Scutarx deltatylus.

Ultrastenos[279]

Gen. et sp. nov

Valid

Stein, Hand & Archer

Late Oligocene

Riversleigh World Heritage Area

 Australia

A member of Mekosuchinae. The type species is U. willisi.

Vivaron[280]

Gen. et sp. nov

Valid

Lessner et al.

Late Triassic (Norian)

Chinle Formation

 United States
( New Mexico)

A rauisuchid. The type species is V. haydeni.

Basal dinosauromorphs

Research[edit]

New taxa[edit]

Name Novelty Status Authors Age Unit Location Notes Images

Dromomeron gigas[284]

Sp. nov

Valid

Martínez et al.

Late Triassic (Norian)

Quebrada del Barro Formation

 Argentina

A lagerpetid dinosauromorph, a species of Dromomeron.

Ixalerpeton[285]

Gen. et sp. nov

Valid

Cabreira et al.

Late Triassic (Carnian)

Santa Maria Formation

 Brazil

A lagerpetid dinosauromorph. The type species is I. polesinensis.

Non-avian dinosaurs

Research[edit]

New taxa[edit]

Name Novelty Status Authors Age Unit Location Notes Images

Agujaceratops mavericus[367]

Sp. nov

Valid[368]

Lehman, Wick & Barnes

Late Cretaceous

Aguja Formation

 United States
( Texas)

A chasmosaurine ceratopsian.

Alcovasaurus[369]

Gen. et comb. nov

Valid

Galton & Carpenter

Late Jurassic

Morrison Formation

 United States
( Wyoming)

A stegosaur; a new genus for "Stegosaurus" longispinus Gilmore (1914). This species was previously made the type species of the new genus Natronasaurus by Ulansky (2014); however, Galton & Carpenter (2016) claim it did not meet the requirements of the International Code of Zoological Nomenclature.[369]

Aoniraptor[370]

Gen. et sp. nov

Valid

Motta et al.

Late Cretaceous (middle Cenomanian-early Turonian)

Huincul Formation

 Argentina

A theropod dinosaur of uncertain phylogenetic placement, a possible relative of Deltadromeus. The type species is A. libertatem.

Apatoraptor[371]

Gen. et sp. nov

Valid

Funston & Currie

Late Cretaceous

Horseshoe Canyon Formation

 Canada
( Alberta)

A caenagnathid theropod. The type species is Apatoraptor pennatus.

Austroposeidon[372]

Gen. et sp. nov

Valid

Bandeira et al.

Late Cretaceous (Campanian-Maastrichtian)

Presidente Prudente Formation

 Brazil

A titanosaur sauropod. The type species is A. magnificus.

Beipiaognathus[373]

Gen. et sp. nov

Valid

Hu, Wang & Huang

Early Cretaceous

Yixian Formation

 China

A compsognathid theropod. The type species is B. jii.

Buriolestes[285]

Gen. et sp. nov

Valid

Cabreira et al.

Late Triassic (Carnian)

Santa Maria Formation

 Brazil

A basal member of Sauropodomorpha. The type species is B. schultzi.

Datonglong[374]

Gen. et sp. nov

Valid

Xu et al.

Late Cretaceous

Huiquanpu Formation

 China

A non-hadrosaurid hadrosauroid ornithopod. The type species is Datonglong tianzhenensis.

Dracoraptor[375]

Gen. et sp. nov

Valid

Martill et al.

Early Jurassic (Hettangian)

Blue Lias Formation

 United Kingdom

A basal member of Neotheropoda. The type species is Dracoraptor hanigani.

Eotrachodon[376][377]

Gen. et sp. nov

Valid

Prieto-Marquez, Erickson & Ebersole

Late Cretaceous (latest Santonian)

Mooreville Chalk

 United States
( Alabama)

A hadrosaurid ornithopod. The type species is Eotrachodon orientalis.

Foraminacephale[378]

Gen. et comb. nov

Valid

Schott & Evans

Late Cretaceous (Campanian)

 Canada
( Alberta)

A new genus for "Stegoceras" brevis Lambe (1918).

Fukuivenator[379]

Gen. et sp. nov

Valid

Azuma et al.

Early Cretaceous (Barremian or Aptian)

Kitadani Formation

 Japan

A member of Maniraptora of uncertain phylogenetic placement. The type species is Fukuivenator paradoxus.

Gastonia lorriemcwhinneyae[380]

Sp. nov

Valid

Kinneer, Carpenter & Shaw

Early Cretaceous

Cedar Mountain Formation

 United States
( Utah)

?Gryposaurus alsatei[381]

Sp. nov

Valid

Lehman, Wick & Wagner

Late Cretaceous (Maastrichtian)

Javelina Formation

 United States
( Texas)

A hadrosaurid, possibly a species of Gryposaurus.

Gualicho[382]

Gen. et sp. nov

Valid

Apesteguía et al.

Late Cretaceous (Cenomanian to Turonian)

Huincul Formation

 Argentina

A theropod dinosaur of uncertain phylogenetic placement, a possible relative of Deltadromeus. The taxon informally referred to as "Nototyrannus" before its formal description. The type species is G. shinyae.

Lohuecotitan[383]

Gen. et sp. nov

Valid

Díaz et al.

Late Cretaceous (late Campanian-early Maastrichtian)

 Spain

A titanosaur sauropod. The type species is L. pandafilandi.

Machairoceratops[384]

Gen. et sp. nov

Valid

Lund et al.

Late Cretaceous (Campanian)

Wahweap Formation

 United States
( Utah)

A centrosaurine ceratopsian. The type species is Machairoceratops cronusi.

Magnamanus[385]

Gen. et sp. nov

Valid

Fuentes Vidarte et al.

Early Cretaceous (late Hauterivian or early Barremian)

Golmayo Formation

 Spain

A basal member of Styracosterna. The type species is M. soriaensis.

Meroktenos[386]

Gen. et comb. nov

Valid

Peyre de Fabrègues & Allain

Late Triassic

Lower Elliot Formation

 Lesotho

A non-sauropod sauropodomorph. The type species is "Melanorosaurus" thabanensis Gauffre (1993).

Morrosaurus[387]

Gen. et sp. nov

Valid

Rozadilla et al.

Late Cretaceous (Maastrichtian)

López de Bertodano Formation

 Antarctica

An iguanodontian ornithopod. The type species is Morrosaurus antarcticus.

Murusraptor[388]

Gen. et sp. nov

Valid

Coria & Currie

Late Cretaceous (Coniacian)

Sierra Barrosa Formation

 Argentina

A theropod belonging to the group Megaraptora. The type species is M. barrosaensis.

Notocolossus[389]

Gen. et sp. nov

Valid

González Riga et al.

Late Cretaceous (late Coniacian–early Santonian)

Plottier Formation

 Argentina

A titanosaur sauropod. The type species is Notocolossus gonzalezparejasi.

Rativates[390]

Gen. et sp. nov

Valid

McFeeters et al.

Late Cretaceous (late Campanian)

Dinosaur Park Formation

 Canada
( Alberta)

An ornithomimid theropod. The type species is R. evadens.

Sarmientosaurus[391]

Gen. et sp. nov

Valid

Martínez et al.

Late Cretaceous (Cenomanian-Turonian)

Bajo Barreal Formation

 Argentina

A titanosaur sauropod, a basal member of Lithostrotia. The type species is Sarmientosaurus musacchioi.

Savannasaurus[392]

Gen. et sp. nov

Poropat et al.

Late Cretaceous (Cenomanian-early Turonian)

Winton Formation

 Australia

A titanosaur sauropod. The type species is S. elliottorum.

Spiclypeus[393]

Gen. et sp. nov

Valid

Mallon et al.

Late Cretaceous (late Campanian)

Judith River Formation

 United States
( Montana)

A chasmosaurine ceratopsian. The type species is Spiclypeus shipporum.

Taurovenator[370]

Gen. et sp. nov

Valid

Motta et al.

Late Cretaceous (middle Cenomanian-early Turonian)

Huincul Formation

 Argentina

A carcharodontosaurid theropod. The type species is T. violantei.

Timurlengia[394]

Gen. et sp. nov

Valid

Brusatte et al.

Late Cretaceous (Turonian)

Bissekty Formation

 Uzbekistan

A non-tyrannosaurid tyrannosauroid. The type species is Timurlengia euotica.

Tongtianlong[395]

Gen. et sp. nov

et al.

Late Cretaceous (Maastrichtian)

Nanxiong Formation

 China

An oviraptorid theropod. The type species is T. limosus.

Tototlmimus[396]

Gen. et sp. nov

Valid

Serrano-Brañas et al.

Late Cretaceous

Packard Shale Formation

 Mexico

An ornithomimid theropod. The type species is Tototlmimus packardensis.

Viavenator[397]

Gen. et sp. nov

Valid

Filippi et al.

Late Cretaceous (Santonian)

Bajo de la Carpa Formation

 Argentina

A brachyrostran abelisaurid theropod. The type species is Viavenator exxoni.

Wiehenvenator [398]

Gen. et sp. nov.

Valid

Rauhut, Hübner & Lanser

Middle Jurassic (Callovian)

Ornatenton Formation

 Germany

A megalosaurid theropod. The type species is W. albati.

Birds

Research[edit]

  • A study on the rates of morphological evolution in Early Cretaceous birds is published by Wang and Lloyd (2016).[399]
  • A study on the microbodies associated with feathers of a new specimen of Eoconfuciusornis from the Early Cretaceous Huajiying Formation (China) and on the matrix in which the microbodies were embedded is published by Pan et al. (2016), who interpret the microbodies as melanosomes.[400]
  • Remains of non-plumage soft tissues, including scales, toe pads, skin and muscle, are identified in two specimens of Confuciusornis by Falk et al. (2016).[401]
  • A skeleton of an enantiornithine bird preserving a gastric pellet that includes fish bones is described from the Early Cretaceous Jehol Biota of China by Wang, Zhou & Sullivan (2016).[402]
  • Two partial wings with vestiges of soft tissues, probably belonging to precocial hatchlings of enantiornithine birds, are described from the Late Cretaceous (Cenomanian) Burmese amber by Xing et al. (2016).[403]
  • A revised diagnosis of Cerebavis cenomanica, a study on the braincase anatomy of the species and a study on its phylogenetic relationships is published by Walsh, Milner & Bourdon (2016).[404]
  • A study on the shape, growth, attachment, implantation, replacement, and tissue microstructures of the teeth of Hesperornis and Ichthyornis is published by Dumont et al. (2016).[405]
  • A phylogenetic analysis of Hesperornithiformes is published by Bell & Chiappe (2016).[406]
  • A specimen of Hesperornis with a healed wound is described from the Late Cretaceous Pierre Shale (South Dakota, United States) by Martin, Rothschild & Burnham (2016), who interpret the wound as caused by an unsuccessful attack of a polycotylid plesiosaur.[407]
  • Pelvic elements of Gargantuavis philoinos, providing new information about the pelvic morphology of the species, are described from the Late Cretaceous (late Campanian/early Maastrichtian) of southern France by Buffetaut & Angst (2016).[408]
  • A specimen of Vegavis iaai with a fossilized syrinx is described from the Late Cretaceous of Antarctica by Clarke et al. (2016).[409]
  • A study on the feeding mechanics and behaviour of five moa species is published by Attard et al. (2016).[410]
  • Mariana B.J. Picasso & María Clelia Mosto, 2016: Hinasuri nehuensis Tambussi was a robust, extinct rheid bird from the early Pliocene of Buenos Aires province, Argentina. This paper revisits the femoral morphology of H. nehuensis and provides an updated osteological description together with new insights into its palaeobiology.[411]
  • Restudies of the Pleistocene species Rhea pampeana and Rhea anchorenensis are published by Picasso (2016) and Picasso and Mosto (2016), respectively, who consider these species to be junior synonyms of the extant greater rhea (Rhea americana).[412][413]
  • Worthy et al. (2016) argue that Sylviornis neocaledoniae is a stem-galliform related to Megavitiornis altirostris and both are placed in the Sylviornithidae Mourer-Chauviré et Balouet, 2005.[414]
  • A revision of the systematics of the early Eocene North American members of Geranoididae is published by Mayr (2016), who argues that geranoidids might be stem group representatives of the Gruoidea (the clade including trumpeters, cranes and related birds).[415]
  • Zelenkov, Boev & Lazaridis (2016) reinterpret Otis hellenica from the Miocene of Greece, originally thought to be a bustard, as a member of Gruiformes belonging to the family Eogruidae and the subfamily Ergilornithinae; the authors classify it as a possible member of the genus Amphipelargus of uncertain specific assignment ("?Amphipelargus sp.").[416]
  • A restudy of the holotype specimen of Bathornis grallator and a study on the taxonomic composition and phylogenetic affinities of bathornithids is published by Mayr (2016).[417]
  • Zelenkov, Volkova and Gorobets (2016) describe buttonquail fossils from the late Miocene of Hungary, southern Ukraine and northern Kazakhstan, and transfer the species Calidris janossyi Kessler (2009) to the genus Ortyxelos.[418]
  • Gerald Mayr and Zbigniew M. Bochenski,(2016) describe a disarticulated postcranial skeleton of a Ralloidea from the Early Oligocene (Rupelian) Jamna Dolna Site 2 in Poland as Gen. et Sp. indet.[419]
  • Agnolin, Tomassini and Contreras (2016) describe a distal end of tarsometatarsus from the late Miocene levels of the Loma de Las Tapias Formation (San Juan Province, Argentina), identified as the oldest seedsnipe fossil discovered so far.[420]
  • Body mass estimates for 25 extinct pan-alcids and a study of body mass evolution in Pan-Alcidae are published by Smith (2016).[421]
  • The earliest known cranial endocast of a stem-penguin (a member of the genus Waimanu) is described from the Paleocene Waipara Greensand (New Zealand) by Proffitt, Clarke & Scofield (2016).[422]
  • Thomas & Ksepka (2016) classify a Whaingaroan penguin from the Glen Massey Formation (North Island, New Zealand), first described in 1973, as a member of the genus Kairuku of uncertain specific assignment, extending the geographic range of the genus.[423]
  • Park et al., 2016 The description of recently collected penguin fossils from the re-dated upper Miocene Port Campbell Limestone of Portland (Victoria), in addition to reanalysis of previously described material, has allowed the Cenozoic history of penguins in Australia to be placed into a global context for the first time. Australian pre-Quaternary fossil penguins represent stem taxa phylogenetically disparate from each other and Eudyptula minor, implying multiple dispersals and extinctions.[424]
  • Carolina Acosta Hospitaleche, Leandro M. Pérez, Sergio Marenssi, Marcelo Reguero (2016). The purpose of this paper is to provide a taphonomic analysis of the holotype of Crossvallia unienwillia, in order to improve the knowledge of the vertebrate record of the Cross Valley Formation, a unit exposed in the central area of Marambio (Seymour) Island, Antarctic Peninsula.[425]
  • A new skeleton of the Eocene penguin Palaeeudyptes klekowskii is described from the Submeseta Formation (Seymour Island, Antarctica) by Acosta Hospitaleche (2016).[426]
  • Carolina Acosta Hospitaleche & Eduardo Olivero, 2016: Eocene penguins are known mostly from Antarctic specimens. A previously documented partial skeleton consisting of a pelvis, femur, tibiotarsus and fibula, from the middle Eocene Leticia Formation, Tierra del Fuego Province, Argentina, has been prepared and re-described. Re-analysis favours assignment to Palaeeudyptes gunnari, a species widely recorded in the Eocene of Antarctica.[427]
  • Fossils of a stork and a heron belonging or related to the tribe Nycticoracini are described from the Pliocene of Myanmar by Stidham et al. (2016).[428]
  • A restudy of the fossils attributed to the species Liornis floweri and Callornis giganteus from the Miocene Santa Cruz Formation (Patagonia, Argentina) is published by Buffetaut (2016), who considers L. floweri to be a junior synonym of Brontornis burmeisteri and considers C. giganteus to be a chimera based on a phorusrhacid tarsometatarsus and a brontornithid tibiotarsus.[429]
  • A study of eggshell fragments from the Pleistocene of Australia putatively referred to Genyornis newtoni is published by Grellet-Tinner, Spooner & Worthy (2016), who argue that these fossils are more likely to be remains of eggs laid by megapodes. Based on the similarities in the structure of eggshells of megapodes and dromornithids, the authors also hypothezise that dromornithids might be a sister group to galliforms rather than to or within anseriforms.[430]
  • A study of burnt putative Genyornis eggshell fragments from the Pleistocene of Australia is published by Miller et al. (2016), who interpret them as confirming that eggs of Genyornis newtoni were harvested by humans.[431]
  • A study on the possible presence, form, and extent of sexual dimorphism in Dromornis stirtoni is published by Handley et al. (2016).[432]
  • Gastornithid and presbyornithid fossils are described from the early Eocene of Ellesmere Island (Canada) by Stidham & Eberle (2016).[433]
  • The genus Wilaru, initially considered to be of a stone-curlew, is reinterpreted as a member of Presbyornithidae by De Pietri et al. (2016); the authors also reassess the Cretaceous species Teviornis gobiensis and confirm it as a member of Presbyornithidae.[434]
  • A revision of anseriform birds known from the late Miocene localities in central Hungary is published by Zelenkov (2016), who transfers the species Anas denesi Kessler (2013) to the genus Aythya and classifies the species Anas albae Janossy (1979) as a member of tribe Mergini of uncertain generic assignment.[435]
  • A revision of galliform birds known from the late Miocene localities in central Hungary is published by Zelenkov (2016), who transfers the subspecies Pavo aesculapi phasianoides Janossy (1991) to the genus Syrmaticus and raises it to the rank of a separate species Syrmaticus phasianoides.[436]
  • New fossil remains of the Eocene cuckoo Chambicuculus pusillus are described from Tunisia by Mourer-Chauviré et al. (2016).[437]
  • Virtual cranial endocast of the dodo is described by Gold, Bourdon & Norell (2016).[438]
  • An ungual phalanx of a large member of Accipitridae belonging to an unknown genus and species is described from the Miocene of Panama by Steadman & MacFadden (2016).[439]
  • Partial tarsometatarsus of a small parrot is described from the Early Miocene Khalagay Formation (Baikal region, Russia) by Zelenkov (2016).[440]
  • Fossil avian feet from the Early Eocene of Messel, Germany are described by Gerald Mayr [441]
  • A new tracksite with bird footprints (attributed to the ichnospecies Uvaichnites riojana), preserved in the early Miocene Lerín Formation (Bardenas Reales de Navarra Natural Park, Navarre, Spain), is described by Díaz-Martínez et al. (2016).[442]
  • A new ichnospecies, Koreananornis lii, from the Lower Cretaceous avian track locality in the Guanshan area, Yongjing County, Gansu Province, northwest China, is described by Xing, Buckley, Lockley, Zhang, Marty, Wang, Li, McCrea et Peng, 2016. (2016).[443]
  • An avian egg from the Lower Cretaceous (Albian) Liangtoutang Formation is described by Lawver et al. (2016) and named Pachycorioolithus jinyunensis oogen. et oosp. nov. within Pachycorioolithidae oofam. nov.[444]
  • Three pellets with bird remains are described from the Eocene Messel pit (Germany) by Mayr & Schaal (2016), who interpret two of the pellets as probably produced by snakes or other squamates, and one as probable owl pellet (which, if confirmed, would make it the oldest owl pellet identified so far), possibly produced by the owl Palaeoglaux artophoron.[445]

New taxa[edit]

Name Novelty Status Authors Age Unit Location Notes Images

Antarctoboenus [446]

Gen. et sp. nov.

Valid

Cenizo, Noriega & Reguero

Early Eocene

La Meseta Formation

 Antarctica

(Seymour Island)

A stem-falconid. The type species is A. carlinii.

Bellulornis [447][448]

Gen. et sp. nov.

Valid

Wang, Zhou & Zhou

Early Cretaceous

Jiufotang Formation

 China

A basal member of Ornithuromorpha. The type species is B. rectusunguis. The original generic name was Bellulia, which turned out to be preoccupied by Bellulia Fibiger (2008).

Calciavis [449]

Gen. et sp. nov.

Valid

Nesbitt & Clarke

Early Eocene

Green River Formation

 United States
( Wyoming)

A member of Lithornithidae. The type species is C. grandei.

Centropus bairdi [450]

Sp. nov.

Valid

Shute, Prideaux & Worthy

Pleistocene

 Australia

A member of the Cuculidae.

Centropus maximus [450]

Sp. nov.

Valid

Shute, Prideaux & Worthy

Pleistocene

 Australia

A member of the Cuculidae.

Changzuiornis [451]

Gen. et sp. nov.

Valid

Huang et al.

Early Cretaceous (Aptian)

Jiufotang Formation

 China

An early member of Euornithes. The type species is C. ahgmi.

Chiappeavis [452]

Gen. et sp. nov.

Valid

O’Connor et al.

Early Cretaceous

Jiufotang Formation

 China

A member of Enantiornithes, probably belonging to the family Pengornithidae. The type species is C. magnapremaxillo.

Chionoides [453]

Gen. et sp. nov.

Valid

De Pietri et al.

Late Oligocene

 Australia

A member of Chionoidea of uncertain phylogenetic placement, showing the mosaic of characters shared with both sheathbills and the Magellanic plover. The type species is C. australiensis.

Chongmingia [454]

Gen. et sp. nov.

Valid

Wang et al.

Early Cretaceous (Aptian)

Jiufotang Formation

 China

A member of Avialae of uncertain phylogenetic placement. The type species is C. zhengi.

Cypseloramphus [455]

Gen. et sp. nov.

Valid

Mayr

Early Eocene

Messel pit

 Germany

Possibly a basal member of Apodiformes. The type species is C. dimidius.

Daphoenositta trevorworthyi [456]

Sp. nov.

Valid

Nguyen

Miocene

Riversleigh World Heritage Area

 Australia

A sittella

Dingavis [457]

Gen. et sp. nov.

Valid [458]

O'Connor, Wang & Hu

Early Cretaceous

Yixian Formation

 China

A basal member of Ornithuromorpha. The type species is D. longimaxilla.

Dromornis murrayi [459]

Sp. nov.

Valid

Worthy et al.

Late Oligocene–Early Miocene

Riversleigh

 Australia

A member of Dromornithidae

Eostrix gulottai [460]

Sp. nov.

Valid

Mayr

Early Eocene

Nanjemoy Formation

 United States
( Virginia)

An early owl of the family Protostrigidae.

Eurobambusicola [436]

Gen. et sp. nov.

Valid

Zelenkov

Late Miocene

 Hungary

A member of the family Phasianidae. The type species is E. turolicus.

Galligeranoides [461]

Gen. et sp. nov.

Valid

Bourdon, Mourer-Chauviré, & Laurent

middle Ypresian

 France

A member of the family Geranoididae. The type species is G. boriensis.

Gallinago kakuki [462]

Sp. nov.

Valid

Steadman & Takano

Late Quaternary

 The Bahamas
 Cayman Islands
 Cuba

A member of Scolopacidae, a species of Gallinago.

Hesperornis lumgairi [463]

Sp. nov.

Valid

Aotsuka & Sato

Campanian

Pierre Shale

 Canada

A species of Hesperornis.

Klallamornis abyssa [464]

Gen. et sp. nov.

Valid

Mayr & Goedert

Latest Eocene or Early Oligocene

 United States
( Washington)

A member of Plotopteridae. This is the type species of the new genus.

?Klallamornis clarki [464]

Sp. nov.

Valid

Mayr & Goedert

Latest Eocene or Early Oligocene

 United States
( Washington)

A member of Plotopteridae. possibly a species of Klallamornis.

Lapillavis [455]

Gen. et sp. nov.

Valid

Mayr

Early Eocene

Messel pit

 Germany

A bird of uncertain phylogenetic placement, showing similarities to Foshanornis songi. The type species is L. incubarens.

Linyiornis [465]

Gen. et sp. nov.

Valid

Wang et al.

Early Cretaceous

Jiufotang Formation

 China

A member of Enantiornithes. The type species is L. amoena.

Mioneophron [466]

Gen. et sp. nov.

Valid

Li et al.

Late Miocene

Liushu Formation

 China

A member of Gypaetinae Vieillot (1816). The type species is M. longirostris.

Mioryaba [436]

Gen. et sp. nov.

Valid

Zelenkov

Late Miocene

 Hungary

A member of the family Phasianidae. The type species is M. magyarica.

Monoenantiornis [467]

Gen. et sp. nov.

Valid[468]

Hu & O’Connor

Early Cretaceous

Yixian Formation

 China

A member of Enantiornithes. The type species is M. sihedangia.

Neilus [453]

Gen. et sp. nov.

Valid

De Pietri et al.

Early Miocene

 New Zealand

A member of Chionoidea of uncertain phylogenetic placement, showing the mosaic of characters shared with both sheathbills and the Magellanic plover. The type species is N. sansomae.

Notoleptos [469]

Gen. et sp. nov

Valid

Acosta Hospitaleche & Gelfo

Late Eocene

 Antarctica

(Seymour Island)

A probable relative of albatrosses. The type species is N. giglii.

Olympidytes [464]

Gen. et sp. nov.

Valid

Mayr & Goedert

Latest Eocene or Early Oligocene

 United States
( Washington)

A member of Plotopteridae. The type species is O. thieli.

Phalcoboenus napieri [470]

Sp. nov.

Valid

Adams & Woods

Holocene

 Falkland Islands

A member of Phalcoboenus.

Primozygodactylus longibrachium [471]

Sp. nov.

Valid

Mayr

Early Eocene

Messel pit

 Germany

A member of Zygodactylidae.

Primozygodactylus quintus [471]

Sp. nov.

Valid

Mayr

Early Eocene

Messel pit

 Germany

A member of Zygodactylidae.

Protomelanitta bakeri [472]

Sp. nov.

Valid

Stidham & Zelenkov

Miocene

Esmeralda Formation

 United States
( Nevada)

A primitive diving duck.

Pseudoseisuropsis wintu [473]

Sp. nov.

Valid

Stefanini, Gómez & Tambussi

Early Pleistocene

Miramar Formation

 Argentina

An ovenbird

Rallus nanus [474]

Nom. nov.

Valid

Alcover et al.

Holocene

 Azores

A member of Rallidae, a species of Rallus; a replacement name for Rallus minutus Alcover et al. (2015) (preoccupied).

Septencoracias [475]

Gen. et sp. nov.

Valid

Bourdon, Kristoffersen & Bonde

Eocene (Ypresian)

Fur Formation

 Denmark

A member of Coracii belonging to the family Primobucconidae. The type species is S. morsensis.

Tingmiatornis [476]

Gen. et sp. nov.

Wang et al.

Late Cretaceous (Turonian)

 Canada
( Nunavut)

A member of Ornithurae of uncertain phylogenetic placement. The type species is T. arctica.

Uria onoi [477]

Sp. nov.

Valid

Watanabe et al.

Late Pleistocene

 Japan

A member of Alcidae

Wilaru prideauxi [434]

Sp. nov.

Valid

De Pietri et al.

Early Miocene

Etadunna Formation
Wipajiri Formation

 Australia

A species of Wilaru.

Pterosaurs

Research[edit]

New taxa[edit]

Name Novelty Status Authors Age Unit Location Notes Images

Allkaruen[482]

Gen. et sp. nov

Valid

Codorniú et al.

Early-Middle Jurassic

Cañadón Asfalto Formation

 Argentina

A non-pterodactyloid member of Breviquartossa. The type species is A. koi.

Aymberedactylus[483]

Gen. et sp. nov

Valid

Pêgas, Leal & Kellner

Early Cretaceous (Aptian-Albian)

Crato Formation

 Brazil

A member of Tapejarinae. The type species is A. cearensis.

Forfexopterus[484]

Gen. et sp. nov

Valid

Jiang et al.

Early Cretaceous

Jiufotang Formation

 China

A member of Archaeopterodactyloidea. The type species is F. jeholensis.

Huaxiapterus atavismus[485]

Sp. nov

Valid

et al.

Early Cretaceous

Jiufotang Formation

 China

Pangupterus[486]

Gen. et sp. nov

Valid

et al.

Early Cretaceous

Jiufotang Formation

 China

A toothed member of Pterodactyloidea. The type species is P. liui.

Sinopterus lingyuanensis[485]

Sp. nov

Valid

et al.

Early Cretaceous

Jiufotang Formation

 China

Other reptiles

Research

  • A skull of a juvenile specimen of Delorhynchus cifellii is described from the Richards Spur locality (Oklahoma, United States) by Haridy et al. (2016).[487]
  • A revision of the systematics of the Chinese pareiasaurs is published by Benton (2016).[488]
  • A study of evolution of body size of the carnivorous and herbivorous members of Captorhinidae is published by Brocklehurst (2016).[489]
  • Surmik et al. (2016) describe nothosaurid and tanystropheid bones from the Triassic of Poland preserving blood-vessel-like structures enclosing organic molecules.[490]
  • Two new specimens of Atopodentatus unicus are described by Chun et al. (2016), providing new information on the skull anatomy of this species and indicating that its rostrum, rather than being downturned as originally assumed, developed a hammerhead-like shape.[491]
  • Description of new material of Hemilopas mentzeli from the Middle Triassic of Silesia (Poland) and a study of the phylogenetic relationships of the species is published by Surmik (2016).[492]
  • Description of the anatomy of partially articulated forelimbs and isolated forelimb bones of Drepanosaurus recovered from the Late Triassic (Norian) Hayden Quarry (Chinle Formation) of New Mexico, USA is published by Pritchard et al. (2016).[493]
  • A study on the femoral and tibial histology of the rhynchosaur Stenaulorhynchus stockleyi is published by Werning & Nesbitt (2016).[494]
  • A study on the maximum body size and distribution of the reptile species known to have gone extinct during the last 50,000 years, as well as the role played by these factors in recent reptile extinction events, is published by Slavenko et al. (2016).[495]

New taxa

Name Novelty Status Authors Age Unit Location Notes Images

Brasinorhynchus[496]

Gen. et sp. nov

Valid

Schultz, Langer & Montefeltro

Middle Triassic (Ladinian)

Santa Maria Formation

 Brazil

A rhynchosaur belonging to the group Stenaulorhynchinae. The type species is Brasinorhynchus mariantensis.

Colobomycter vaughni[497]

Sp. nov

Valid

MacDougall, Modesto & Reisz

Early Permian

 United States
( Oklahoma)

A member of Lanthanosuchoidea.

Euconcordia[498]

Nom. nov

Valid

Reisz, Haridy & Müller

Carboniferous (Pennsylvanian)

Calhouns Shale

 United States
( Kansas)

A member of Captorhinidae; a replacement name for Concordia Müller & Reisz (2005).

Langeronyx[499]

Gen. et comb. nov

Valid

Ezcurra, Montefeltro & Butler

Middle Triassic (Anisian)

Bromsgrove Sandstone Formation

 United Kingdom

A rhynchosaur; a new genus for "Rhynchosaurus" brodiei Benton (1990).

Ozimek[500]

Gen. et sp. nov

Valid

Dzik & Sulej

Late Triassic (probably late Carnian)

 Poland

A relative of Sharovipteryx. The type species is O. volans.

Teyujagua[501]

Gen. et sp. nov

Valid

Pinheiro et al.

Early Triassic (Induan to early Olenekian)

Sanga do Cabral Formation

 Brazil

A member of Archosauromorpha closely related to Archosauriformes. The type species is Teyujagua paradoxa.

Xinpusaurus xingyiensis[502]

Sp. nov

Valid

Li et al.

Middle Triassic (Ladinian)

 China

A thalattosaur.

Synapsids

Non-mammalian synapsids

Research[edit]

New taxa[edit]

Name Novelty Status Authors Age Unit Location Notes Images

Abdalodon[515]

Gen. et sp. nov

Valid

Kammerer

Late Permian

Beaufort Group
(Tropidostoma Assemblage Zone)

 South Africa

An early cynodont related to Charassognathus gracilis. The type species is Abdalodon diastematicus.

Bonacynodon[516]

Gen. et sp. nov

Valid

Martinelli, Soares & Schwanke

Late Triassic (early Carnian)

Santa Maria Supersequence

 Brazil

A member of Probainognathidae. The type species is B. schultzi.

Montirictus[517]

Gen. et sp. nov

Valid

Matsuoka, Kusuhashi & Korfe

Early Cretaceous (probably Barremian or Aptian)

Kuwajima Formation

 Japan

A member of Tritylodontidae. The type species is Montirictus kuwajimaensis. Averianov et al. (2017) considered the genus Montirictus to be a junior synonym of the genus Stereognathus.[518]

Mupashi[519]

Gen. et sp. nov

Valid

Huttenlocker & Sidor

Late Permian

Madumabisa Mudstone Formation

 Zambia

A therocephalian related to Karenites. The type species is Mupashi migrator.

Rastodon[520]

Gen. et sp. nov

Valid

Boos et al.

Permian (Guadalupian/Lopingian)

Rio do Rasto Formation

 Brazil

A dicynodont belonging to the group Bidentalia. The type species is R. procurvidens.

Santacruzgnathus[516]

Gen. et sp. nov

Valid

Martinelli, Soares & Schwanke

Late Triassic (early Carnian)

Santa Maria Supersequence

 Brazil

A member of Probainognathia, probably closely related to prozostrodontians. The type species is S. abdalai.

Sauroscaptor[521]

Gen. et sp. nov

Valid

Kammerer, Bandyopadhyay & Ray

Late Permian

Kundaram Formation

 India

A cistecephalid dicynodont. The type species is S. tharavati.

Vaughnictis[522]

Gen. et comb. nov

Valid

Brocklehurst et al.

Permian (Asselian-Sakmarian)

Cutler Group

 United States
( Colorado)

A member of Eothyrididae; a new genus for "Mycterosaurus" smithae Lewis & Vaughn (1965).

Wantulignathus[523]

Gen. et sp. nov

Valid

Whitney & Sidor

Permian (Guadalupian)

Madumabisa Mudstone Formation

 Zambia

A biarmosuchian. The type species is Wantulignathus gwembensis.

Mammals

Other animals

Research

New taxa

Name Novelty Status Authors Age Unit Location Notes Images

Acanthograptus lateralis[531]

Sp. nov

Valid

Chen in Chen et al.

Ordovician

 China

A graptolite.

Acoelia norica[532]

Sp. nov

Valid

Senowbari-Daryan & Link

Late Triassic (Norian)

 Turkey

A demosponge belonging to the group Agelasida.

Acrograptus tenuiculus[531]

Sp. nov

Valid

Chen in Chen et al.

Ordovician

 China

A graptolite.

Amblysiphonella aiyongcuoensis[533]

Sp. nov

Valid

Deng

Permian

 China

A sponge.

Annulitubus[534]

Gen. et sp. nov

Valid

Vinn et al.

Devonian (late Emsian)

Ponta Grossa Formation

 Brazil

An annelid, possibly a polychaete. The type species is Annulitubus mutveii.

Antennipatus[535]

Gen. et sp. nov

Valid

Garwood, Edgecombe & Giribet in Garwood et al.

Carboniferous (Stephanian)

Montceau-les-Mines Lagerstätte

 France

A velvet worm. The type species is A. montceauensis.

Apoglossograptus uniformis[531]

Sp. nov

Valid

Chen in Chen et al.

Ordovician

 China

A graptolite.

Archiasterella dhiraji[536]

Sp. nov

Valid

Gilbert, Hughes & Myrow

Cambrian

Parahio Formation

 India

Archiclimacograptus columnus[531]

Sp. nov

Valid

Chen in Chen et al.

Ordovician

 China

A graptolite.

Baltichaeta[537]

Gen. et sp. nov

Valid[538]

Slater et al.

Cambrian (Stage 4)

File Haidar Formation

 Sweden

A member of the total group of Annelida. The type species is B. jormunganda.

Baltiscalida[537]

Gen. et sp. nov

Valid[538]

Slater et al.

Cambrian (Stage 4)

File Haidar Formation

 Sweden

A member of the total group of Priapulida. The type species is B. njorda.

Bistella[539]

Gen. et sp. nov

Valid

Fedorov in Fedorov, Parkhaev & Demidenko

Cambrian

 Russia

A heteractinid sponge belonging to the group Heteractinellida and the family Heterostellidae. The type species is B. inexplicabila.

Bohemograptus praecox[540]

Sp. nov

Valid

Štorch et al.

Silurian

 Czech Republic

A graptolite.

Bolidium bertii[541]

Sp. nov

Valid[542]

Frisone, Pisera & Preto

Eocene (early Lutetian)

 Italy

A sponge, a species of Bolidium.

Burejospermum punctatum[543]

Sp. nov

Valid

McLoughlin et al.

Early Eocene

La Meseta Formation

Antarctica
(Seymour Island)

A member of Clitellata of uncertain phylogenetic placement, described on the basis of fossilized cocoons; a species of Burejospermum.

Burejospermum seymourense[543]

Sp. nov

Valid

McLoughlin et al.

Early Eocene

La Meseta Formation

Antarctica
(Seymour Island)

A member of Clitellata of uncertain phylogenetic placement, described on the basis of fossilized cocoons; a species of Burejospermum.

Caelispongia[532]

Gen. et sp. nov

Valid

Senowbari-Daryan & Link

Late Triassic (Norian)

Kasımlar Basin

 Turkey

A demosponge belonging to the group Agelasida and the family Stellispongiellidae. The type species is C. topukensis.

Camerospongia tuberculata[541]

Sp. nov

Valid[542]

Frisone, Pisera & Preto

Eocene (early Lutetian)

 Italy

A sponge, a species of Camerospongia.

Camerospongia visentinae[541]

Sp. nov

Valid[542]

Frisone, Pisera & Preto

Eocene (early Lutetian)

 Italy

A sponge, a species of Camerospongia.

Capsulocyathus petri[544]

Sp. nov

Valid

Sundukov & Zhuravlev

Cambrian

 Russia

A member of Archaeocyatha belonging to the family Cryptoporocyathidae.

Cavispongia scarpai[541]

Sp. nov

Valid[542]

Frisone, Pisera & Preto

Eocene (early Lutetian)

 Italy

A sponge, a species of Cavispongia.

Conotheca hensoni[545]

Sp. nov

Valid

Peel et al.

Cambrian

Henson Gletscher Formation

 Greenland

A member of Hyolitha belonging to the group Orthothecida.

Constellatispongia[546]

Gen. et sp. nov

Valid

Botting & Peel

Early Cambrian

Buen Formation

 Greenland

A sponge. The type species is C. canismajorii.

Corallistes multiosculata[541]

Sp. nov

Valid[542]

Frisone, Pisera & Preto

Eocene (early Lutetian)

 Italy

A sponge, a species of Corallistes.

Cornulites vilcae[547]

Sp. nov

Valid

Vinn & Gutiérez-Marco

Late Ordovician (probably late Sandbian)

Probably Calapuja Formation

 Peru

A member of Cornulitida (a group of animals of uncertain phylogenetic placement, possibly molluscs), a species of Cornulites.

Cornulites zatoni[547]

Sp. nov

Valid

Vinn & Gutiérez-Marco

Late Ordovician (Sandbian)

Calapuja Formation

 Peru

A member of Cornulitida (a group of animals of uncertain phylogenetic placement, possibly molluscs), a species of Cornulites.

Coronispongia[541]

Gen. et sp. nov

Valid[542]

Frisone, Pisera & Preto

Eocene (early Lutetian)

 Italy

A sponge. The type species is Coronispongia confossa.

Corynites nanus[531]

Sp. nov

Valid

Chen in Chen et al.

Ordovician

 China

A graptolite.

Crassicoactum[546]

Gen. et sp. nov

Valid

Botting & Peel

Early Cambrian

Buen Formation

 Greenland

A sponge. The type species is C. cucumis.

Daharella triassica[532]

Sp. nov

Valid

Senowbari-Daryan & Link

Late Triassic (Norian)

Kasımlar Basin

 Turkey

A demosponge belonging to the group Agelasida and the family Auriculospongiidae.

Dicranograptus ramosus angustus[531]

Subsp. nov

Valid

Chen in Chen et al.

Ordovician

 China

A graptolite.

Eoglyptograptus asymmetros[531]

Sp. nov

Valid

Goldman & Zhang in Chen et al.

Ordovician

 China

A graptolite.

Eolipastrotethya[548]

Gen. et sp. nov

Valid[549]

Łukowiak & Pisera

Late Eocene

Pallinup Formation

 Australia

A heteroscleromorph demosponge belonging to the group Bubarida. The type species is E. picketti.

Erismacoscinus lucanoi[550]

Sp. nov

Valid

Menéndez et al.

Cambrian Stage 3

Pedroche Formation

 Spain

A member of Archaeocyatha belonging to the family Asterocyathidae.

Esakovella[539]

Gen. et sp. nov

Valid

Fedorov in Fedorov, Parkhaev & Demidenko

Cambrian

 Russia

A tommotiid belonging to the family Kelanellidae. The type species is E. grigorievae.

Fallocyathus accomodatus[544]

Sp. nov

Valid

Sundukov in Sundukov & Zhuravlev

Cambrian

 Russia

A member of Archaeocyatha belonging to the family Fallocyathidae.

Fallocyathus apheles[544]

Sp. nov

Valid

Sundukov in Sundukov & Zhuravlev

Cambrian

 Russia

A member of Archaeocyatha belonging to the family Fallocyathidae.

Geodia hopetouni[548]

Sp. nov

Valid[549]

Łukowiak & Pisera

Late Eocene

Pallinup Formation

 Australia

A heteroscleromorph demosponge belonging to the group Tetractinellida, a species of Geodia.

Haddingograptus cuneatus[531]

Sp. nov

Valid

Chen in Chen et al.

Ordovician

 China

A graptolite.

Haddingograptus flexibilis[531]

Sp. nov

Valid

Chen in Chen et al.

Ordovician

 China

A graptolite.

Haddingograptus tarimensis[531]

Sp. nov

Valid

Chen in Chen et al.

Ordovician

 China

A graptolite.

Hadimopanella incubo[551]

Sp. nov

Valid[552]

Streng, Ebbestad & Berg-Madsen

Cambrian

 Sweden

A palaeoscolecid worm.

Hadimopanella oelandiana[551]

Sp. nov

Valid[552]

Streng, Ebbestad & Berg-Madsen

Cambrian

 Sweden

A palaeoscolecid worm.

Hallograptus echinatus[531]

Sp. nov

Valid

Chen in Chen et al.

Ordovician

 China

A graptolite.

Hamptonia limatula[546]

Sp. nov

Valid

Botting & Peel

Early Cambrian

Buen Formation

 Greenland

A sponge.

Hexactinella clampensis[541]

Sp. nov

Valid[542]

Frisone, Pisera & Preto

Eocene (early Lutetian)

 Italy

A sponge, a species of Hexactinella.

?Hyalostelia spinula[539]

Sp. nov

Valid

Fedorov in Fedorov, Parkhaev & Demidenko

Cambrian

 Russia

A hexactinellid sponge belonging to the group Hexasterophora and the family Hyalostellidae.

Jiangxigraptus? delicatus[531]

Sp. nov

Valid

Chen in Chen et al.

Ordovician

 China

A graptolite.

Jiangxigraptus ultilis[531]

Sp. nov

Valid

Chen in Chen et al.

Ordovician

 China

A graptolite.

Kuonamia[553]

Nom. nov

Valid

Doweld

Cambrian Stage 4 to Guzhangian

Holm Dal Formation
Kuonamka Formation

 Greenland[554]
 Russia
( Sakha Republic)

A sponge; a replacement name for Disparella Fedorov in Fedorov & Pereladov (1987).

Lenica perversa[546]

Sp. nov

Valid

Botting & Peel

Early Cambrian

Buen Formation

 Greenland

A sponge.

Lyrarapax trilobus[555]

Sp. nov

Valid

Cong et al.

Early Cambrian

 China

Milyasa[532]

Gen. et 2 sp. nov

Valid

Senowbari-Daryan & Link

Late Triassic (Norian)

Dereköy Basin

 Turkey

A demosponge belonging to the group Agelasida and the family Preperonidellidae. The type species is M. polysiphonata; genus also includes M. askomorpha.

Mimograptus tenuis[531]

Sp. nov

Valid

Chen in Chen et al.

Ordovician

 China

A graptolite.

Monocrepidium pauli[548]

Sp. nov

Valid[549]

Łukowiak & Pisera

Late Eocene

Pallinup Formation

 Australia

A heteroscleromorph demosponge belonging to the group Bubarida.

Monocrepidium unispiculatum[548]

Sp. nov

Valid[549]

Łukowiak & Pisera

Late Eocene

Pallinup Formation

 Australia

A heteroscleromorph demosponge belonging to the group Bubarida.

Nganki namak[556]

Sp. nov

Valid

Kruse & Hughes

Cambrian

Khussak Formation

 Pakistan

A member of Hyolitha (a group of animals of uncertain phylogenetic placement, possibly molluscs), a species of Nganki.

Obscurospongia[532]

Gen. et sp. nov

Valid

Senowbari-Daryan & Link

Late Triassic (Norian)

Kasımlar Basin

 Turkey

An inozoid or chaetetid sponge. The type species is O. chaetetiformis

Oepikograptus originalis[531]

Sp. nov

Valid

Chen in Chen et al.

Ordovician

 China

A graptolite.

Orthograptus paracalcaratus[531]

Sp. nov

Valid

Chen in Chen et al.

Ordovician

 China

A graptolite.

Ottoia guizhouensis[557]

Sp. nov

Valid

Yang, Zhao & Zhang

Cambrian

Kaili Formation

 China

A stem-group priapulid, a species of Ottoia.

Pachastrella australis[548]

Sp. nov

Valid[549]

Łukowiak & Pisera

Late Eocene

Pallinup Formation

 Australia

A heteroscleromorph demosponge belonging to the group Tetractinellida.

Pachastrella intermedia[548]

Sp. nov

Valid[549]

Łukowiak & Pisera

Late Eocene

Pallinup Formation

 Australia

A heteroscleromorph demosponge belonging to the group Tetractinellida.

Palaeoscolex xinglongensis[558]

Sp. nov

Valid

Liu et al.

Cambrian

Wulongqing Formation

 China

A palaeoscolecid.

Palaeotubus[559]

Gen. et sp. nov

Valid

Sanfilippo et al.

Permian

Pietra di Salomone limestone

 Italy

A polychaete, possibly a relative of serpulids. The type species is P. sosiensis.

Pegmatothylakos[543]

Gen. et sp. nov

Valid

McLoughlin et al.

Early Eocene

La Meseta Formation

Antarctica
(Seymour Island)

A member of Clitellata of uncertain phylogenetic placement, described on the basis of fossilized cocoons. The type species is Pegmatothylakos manumii.

Phycopsis arbusculum[548]

Sp. nov

Valid[549]

Łukowiak & Pisera

Late Eocene

Pallinup Formation

 Australia

A heteroscleromorph demosponge belonging to the group Axinellida.

Preperonidella asymmetrica[532]

Sp. nov

Valid

Senowbari-Daryan & Link

Late Triassic (Norian)

Kasımlar Basin

 Turkey

A demosponge belonging to the group Agelasida and the family Preperonidellidae.

Proclimacograptus angustatus ultimus[531]

Subsp. nov

Valid

Chen in Chen et al.

Ordovician

 China

A graptolite.

Pronormalograptus[531]

Gen. et 2 sp. et comb. nov

Valid

Chen in Chen et al.

Ordovician

 China
 United Kingdom

A graptolite. The type species is P. acicularis; genus also includes new species P. regularis, as well as P. euglyphus (Lapworth, 1880) and P. siccatus (Elles and Wood, 1907).

Pseudosardospongia[539]

Gen. et 2 sp. nov

Valid

Fedorov in Fedorov, Parkhaev & Demidenko

Cambrian

 Russia

A calcareous sponge belonging to the family Polyactinellidae. The type species is P. plana; genus also includes P. miriculata.

Rajatubulus[560]

Gen. et comb. nov

Valid

Yang et al.

Early Cambrian

 Kazakhstan

A member of Cloudinidae; a new genus for "Pseudorthotheca" costata Mambetov in Missarzhevsky & Mambetov (1981).

Rankenella zhangxianensis[561]

Sp. nov

Valid

Lee, Woo & Lee

Cambrian

Zhangxia Formation

 China

A sponge belonging to the family Anthaspidellidae.

Ratcliffespongia freuchenensis[546]

Sp. nov

Valid

Botting & Peel

Early Cambrian

Buen Formation

 Greenland

A sponge.

Rigonia[541]

Gen. et sp. nov

Valid[542]

Frisone, Pisera & Preto

Eocene (early Lutetian)

 Italy

A sponge. The type species is Rigonia plicata.

Rotundocyathus indistinctus[544]

Sp. nov

Valid

Sundukov in Sundukov & Zhuravlev

Cambrian

 Russia

A member of Archaeocyatha belonging to the family Ajacicyathidae.

Rugatotheca daibuica[560]

Sp. nov

Valid

Yang et al.

Early Cambrian

Daibu Member of the Xiaotan section

 China

A small shelly fossil of uncertain phylogenetic placement.

Saccoglossus testa[562]

Sp. nov

Valid

Cameron

Carboniferous (Pennsylvanian)

Mazon Creek fossil beds

 United States
( Illinois)

An acorn worm belonging to the family Harrimaniidae, a species of Saccoglossus.

Saetaspongia procera[546]

Sp. nov

Valid

Botting & Peel

Early Cambrian

Buen Formation

 Greenland

A sponge.

Sanxiascolex[563]

Gen. et sp. nov

Valid

Yang & Zhang

Cambrian

Shipai Formation

 China

A palaeoscolecid. The type species is P. papillogyrus.

Sardospongia gigantea[539]

Sp. nov

Valid

Fedorov in Fedorov, Parkhaev & Demidenko

Cambrian

 Russia

A calcareous sponge belonging to the family Polyactinellidae.

Sardospongia triplexa[539]

Sp. nov

Valid

Fedorov in Fedorov, Parkhaev & Demidenko

Cambrian

 Russia

A calcareous sponge belonging to the family Polyactinellidae.

Sarmentofascis zamparelliae[564]

Sp. nov

Valid

Schlagintweit, Frijia & Parente

Late Cretaceous (early Campanian)

 Italy

A sponge, a species of Sarmentofascis.

Semigothograptus[565]

Gen. et comb. nov

Valid

Kozłowska

Silurian

 Czech Republic
 Germany
 Poland
 United Kingdom

A graptolite; a new genus for "Gothograptus" meganassa Rickards & Palmer (2002).

Sibirecyathus onkhoydokh[544]

Sp. nov

Valid

Sundukov & Zhuravlev

Cambrian

 Russia

A member of Archaeocyatha belonging to the family Ajacicyathidae.

Spathioconchus[566]

Gen. et sp. nov

Zatoń et al.

Early Triassic (Induan)

 Greenland

A member of Microconchida (a group of animals of uncertain phylogenetic placement, possibly molluscs). The type species is S. weedoni.

Spinuliconchus[567]

Gen. et et comb. sp. nov

Valid

Zatoń & Olempska

Devonian

 Poland
 United States

A member of Microconchida. The type species is "Spirorbis" angulatus Hall (1861); genus also includes new species S. biernatae.

Stauractinella eocenica[541]

Sp. nov

Valid[542]

Frisone, Pisera & Preto

Eocene (early Lutetian)

 Italy

A sponge, a species of Stauractinella.

Talacastospongia[568]

Gen. et sp. nov

Valid

Carrera & Rustán

Devonian (Lochkovian)

Talacasto Formation

 Argentina

A hexactinellid sponge. The type species is Talacastospongia minima.

Tanchajella[539]

Gen. et sp. nov

Valid

Fedorov in Fedorov, Parkhaev & Demidenko

Cambrian

 Russia

A hexactinellid sponge belonging to the group Amphidiscophora and the family Hyalonematidae. The type species is T. aculeata.

Taurispongia[532]

Gen. et 8 sp. nov

Valid

Senowbari-Daryan & Link

Late Triassic (Norian)

Dereköy Basin

 Turkey

A demosponge belonging to the group Agelasida and the family Stellispongiellidae. The type species is T. oligocanalis; genus also includes new species T. polycanalis, T. tenuis, T. lamellicanalis, T. fascifera, T. siderifera, T. lamellata and T. polyforma.

Tenuipariespongia taurica[532]

Sp. nov

Valid

Senowbari-Daryan & Link

Late Triassic (Norian)

Kasımlar Basin

 Turkey

A demosponge belonging to the group Agelasida.

Toulminia italica[541]

Sp. nov

Valid[542]

Frisone, Pisera & Preto

Eocene (early Lutetian)

 Italy

A sponge, a species of Toulminia.

Triptolemma solida[548]

Sp. nov

Valid[549]

Łukowiak & Pisera

Late Eocene

Pallinup Formation

 Australia

A heteroscleromorph demosponge belonging to the group Tetractinellida.

Tritonychus [569]

Gen. et sp. nov.

Valid

Zhang et al.

Early Cambrian

 China

A lobopodian. The type species is T. phanerosarkus.

Tuberoconchus[567]

Gen. et sp. nov

Valid

Zatoń in Zatoń & Olempska

Silurian (Ludlow)

Kuressaare Formation